{"bundle_type":"pith_open_graph_bundle","bundle_version":"1.0","pith_number":"pith:2019:2OBTYOIM4TTAL3H4A2SPOJQPV7","short_pith_number":"pith:2OBTYOIM","canonical_record":{"source":{"id":"1910.14645","kind":"arxiv","version":1},"metadata":{"license":"http://arxiv.org/licenses/nonexclusive-distrib/1.0/","primary_cat":"physics.bio-ph","submitted_at":"2019-10-31T17:35:00Z","cross_cats_sorted":["cond-mat.soft"],"title_canon_sha256":"be9a83831eaca612dfca2ff729e8dcc4c70c08a6ec9b60a420dc885e1f77486d","abstract_canon_sha256":"7accdaa4e3a1f05205aa7131a24d51ed49576e1274967b66bc344238cbf46511"},"schema_version":"1.0"},"canonical_sha256":"d3833c390ce4e605ecfc06a4f7260fafc661d04fae1045c3f256d0d3d5e10dcb","source":{"kind":"arxiv","id":"1910.14645","version":1},"source_aliases":[{"alias_kind":"arxiv","alias_value":"1910.14645","created_at":"2026-07-05T00:59:54Z"},{"alias_kind":"arxiv_version","alias_value":"1910.14645v1","created_at":"2026-07-05T00:59:54Z"},{"alias_kind":"doi","alias_value":"10.48550/arxiv.1910.14645","created_at":"2026-07-05T00:59:54Z"},{"alias_kind":"pith_short_12","alias_value":"2OBTYOIM4TTA","created_at":"2026-07-05T00:59:54Z"},{"alias_kind":"pith_short_16","alias_value":"2OBTYOIM4TTAL3H4","created_at":"2026-07-05T00:59:54Z"},{"alias_kind":"pith_short_8","alias_value":"2OBTYOIM","created_at":"2026-07-05T00:59:54Z"}],"events":[{"event_type":"record_created","subject_pith_number":"pith:2019:2OBTYOIM4TTAL3H4A2SPOJQPV7","target":"record","payload":{"canonical_record":{"source":{"id":"1910.14645","kind":"arxiv","version":1},"metadata":{"license":"http://arxiv.org/licenses/nonexclusive-distrib/1.0/","primary_cat":"physics.bio-ph","submitted_at":"2019-10-31T17:35:00Z","cross_cats_sorted":["cond-mat.soft"],"title_canon_sha256":"be9a83831eaca612dfca2ff729e8dcc4c70c08a6ec9b60a420dc885e1f77486d","abstract_canon_sha256":"7accdaa4e3a1f05205aa7131a24d51ed49576e1274967b66bc344238cbf46511"},"schema_version":"1.0"},"canonical_sha256":"d3833c390ce4e605ecfc06a4f7260fafc661d04fae1045c3f256d0d3d5e10dcb","receipt":{"kind":"pith_receipt","key_id":"pith-v1-2026-05","algorithm":"ed25519","signed_at":"2026-07-05T00:59:54.972626Z","signature_b64":"xSd+rHkTX8M1XOrU6I1+ZMxw+eX6gzewLC8vBMQ6dyB0uynmEtRTWFdF+5TBjWXdqX6ZO7UENN3QHh0RZg9DDQ==","signed_message":"canonical_sha256_bytes","builder_version":"pith-number-builder-2026-05-17-v1","receipt_version":"0.3","canonical_sha256":"d3833c390ce4e605ecfc06a4f7260fafc661d04fae1045c3f256d0d3d5e10dcb","last_reissued_at":"2026-07-05T00:59:54.972092Z","signature_status":"signed_v1","first_computed_at":"2026-07-05T00:59:54.972092Z","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54"},"source_kind":"arxiv","source_id":"1910.14645","source_version":1,"attestation_state":"computed"},"signer":{"signer_id":"pith.science","signer_type":"pith_registry","key_id":"pith-v1-2026-05","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54"},"created_at":"2026-07-05T00:59:54Z","supersedes":[],"prev_event":null,"signature":{"signature_status":"signed_v1","algorithm":"ed25519","key_id":"pith-v1-2026-05","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54","signature_b64":"WjYCqKFNirleSUiM1Gsu9K7M2YrMyPhx98QjmK4NmZgQ+N5LzuLJDoLDKMDU3gEaZw8AbwTGpSQUIl4pjeGFCg==","signed_message":"open_graph_event_sha256_bytes","signed_at":"2026-08-04T00:58:32.961216Z"},"content_sha256":"d4e3e9edf6eb23a472adc40e0a88579e4338ce1faa32d842d31339d56e494846","schema_version":"1.0","event_id":"sha256:d4e3e9edf6eb23a472adc40e0a88579e4338ce1faa32d842d31339d56e494846"},{"event_type":"graph_snapshot","subject_pith_number":"pith:2019:2OBTYOIM4TTAL3H4A2SPOJQPV7","target":"graph","payload":{"graph_snapshot":{"paper":{"title":"Non-uniform growth and surface friction determine bacterial biofilm morphology on soft substrates","license":"http://arxiv.org/licenses/nonexclusive-distrib/1.0/","headline":"","cross_cats":["cond-mat.soft"],"primary_cat":"physics.bio-ph","authors_text":"Andrej Kosmrlj, Bonnie L. Bassler, Chenyi Fei, Howard A. Stone, Jing Yan, Ned S. Wingreen, Ricard Alert, Sheng Mao","submitted_at":"2019-10-31T17:35:00Z","abstract_excerpt":"During development, organisms acquire three-dimensional shapes with important physiological consequences. While the basic mechanisms underlying morphogenesis are known in eukaryotes, it is often difficult to manipulate them in vivo. To circumvent this issue, here we present a study of developing Vibrio cholerae biofilms grown on agar substrates in which the spatiotemporal morphological patterns were altered by varying the agar concentration. Expanding biofilms are initially flat, but later experience a mechanical instability and become wrinkled. Whereas the peripheral region develops ordered r"},"claims":{"count":0,"items":[],"snapshot_sha256":"258153158e38e3291e3d48162225fcdb2d5a3ed65a07baac614ab91432fd4f57"},"source":{"id":"1910.14645","kind":"arxiv","version":1},"verdict":{"id":null,"model_set":{},"created_at":null,"strongest_claim":"","one_line_summary":"","pipeline_version":null,"weakest_assumption":"","pith_extraction_headline":""},"integrity":{"clean":true,"summary":{"advisory":0,"critical":0,"by_detector":{},"informational":0},"endpoint":"/pith/1910.14645/integrity.json","findings":[],"available":true,"detectors_run":[],"snapshot_sha256":"c28c3603d3b5d939e8dc4c7e95fa8dfce3d595e45f758748cecf8e644a296938"},"references":{"count":0,"sample":[],"resolved_work":0,"snapshot_sha256":"258153158e38e3291e3d48162225fcdb2d5a3ed65a07baac614ab91432fd4f57","internal_anchors":0},"formal_canon":{"evidence_count":0,"snapshot_sha256":"258153158e38e3291e3d48162225fcdb2d5a3ed65a07baac614ab91432fd4f57"},"author_claims":{"count":0,"strong_count":0,"snapshot_sha256":"258153158e38e3291e3d48162225fcdb2d5a3ed65a07baac614ab91432fd4f57"},"builder_version":"pith-number-builder-2026-05-17-v1"},"verdict_id":null},"signer":{"signer_id":"pith.science","signer_type":"pith_registry","key_id":"pith-v1-2026-05","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54"},"created_at":"2026-07-05T00:59:54Z","supersedes":[],"prev_event":null,"signature":{"signature_status":"signed_v1","algorithm":"ed25519","key_id":"pith-v1-2026-05","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54","signature_b64":"R1XlwK5JFSTOOz7bu+Mw9tZ/TbdnyqS3jN6mtZ1ZleePHjY9tvgJex3yhk+TWzfmHGkyEJeS33jjCGyDwG+2DQ==","signed_message":"open_graph_event_sha256_bytes","signed_at":"2026-08-04T00:58:32.961712Z"},"content_sha256":"7256c4f993747898f657645393b220f6aab1d483881d6f381b85b94e2b5156f1","schema_version":"1.0","event_id":"sha256:7256c4f993747898f657645393b220f6aab1d483881d6f381b85b94e2b5156f1"}],"timestamp_proofs":[],"mirror_hints":[{"mirror_type":"https","name":"Pith Resolver","base_url":"https://pith.science","bundle_url":"https://pith.science/pith/2OBTYOIM4TTAL3H4A2SPOJQPV7/bundle.json","state_url":"https://pith.science/pith/2OBTYOIM4TTAL3H4A2SPOJQPV7/state.json","well_known_bundle_url":"https://pith.science/.well-known/pith/2OBTYOIM4TTAL3H4A2SPOJQPV7/bundle.json","status":"primary"}],"public_keys":[{"key_id":"pith-v1-2026-05","algorithm":"ed25519","format":"raw","public_key_b64":"stVStoiQhXFxp4s2pdzPNoqVNBMojDU/fJ2db5S3CbM=","public_key_hex":"b2d552b68890857171a78b36a5dccf368a953413288c353f7c9d9d6f94b709b3","fingerprint_sha256_b32_first128bits":"RVFV5Z2OI2J3ZUO7ERDEBCYNKS","fingerprint_sha256_hex":"8d4b5ee74e4693bcd1df2446408b0d54","rotates_at":null,"url":"https://pith.science/pith-signing-key.json","notes":"Pith uses this Ed25519 key to sign canonical record SHA-256 digests. Verify with: ed25519_verify(public_key, message=canonical_sha256_bytes, signature=base64decode(signature_b64))."}],"merge_version":"pith-open-graph-merge-v1","built_at":"2026-08-04T00:58:32Z","links":{"resolver":"https://pith.science/pith/2OBTYOIM4TTAL3H4A2SPOJQPV7","bundle":"https://pith.science/pith/2OBTYOIM4TTAL3H4A2SPOJQPV7/bundle.json","state":"https://pith.science/pith/2OBTYOIM4TTAL3H4A2SPOJQPV7/state.json","well_known_bundle":"https://pith.science/.well-known/pith/2OBTYOIM4TTAL3H4A2SPOJQPV7/bundle.json"},"state":{"state_type":"pith_open_graph_state","state_version":"1.0","pith_number":"pith:2019:2OBTYOIM4TTAL3H4A2SPOJQPV7","merge_version":"pith-open-graph-merge-v1","event_count":2,"valid_event_count":2,"invalid_event_count":0,"equivocation_count":0,"current":{"canonical_record":{"metadata":{"abstract_canon_sha256":"7accdaa4e3a1f05205aa7131a24d51ed49576e1274967b66bc344238cbf46511","cross_cats_sorted":["cond-mat.soft"],"license":"http://arxiv.org/licenses/nonexclusive-distrib/1.0/","primary_cat":"physics.bio-ph","submitted_at":"2019-10-31T17:35:00Z","title_canon_sha256":"be9a83831eaca612dfca2ff729e8dcc4c70c08a6ec9b60a420dc885e1f77486d"},"schema_version":"1.0","source":{"id":"1910.14645","kind":"arxiv","version":1}},"source_aliases":[{"alias_kind":"arxiv","alias_value":"1910.14645","created_at":"2026-07-05T00:59:54Z"},{"alias_kind":"arxiv_version","alias_value":"1910.14645v1","created_at":"2026-07-05T00:59:54Z"},{"alias_kind":"doi","alias_value":"10.48550/arxiv.1910.14645","created_at":"2026-07-05T00:59:54Z"},{"alias_kind":"pith_short_12","alias_value":"2OBTYOIM4TTA","created_at":"2026-07-05T00:59:54Z"},{"alias_kind":"pith_short_16","alias_value":"2OBTYOIM4TTAL3H4","created_at":"2026-07-05T00:59:54Z"},{"alias_kind":"pith_short_8","alias_value":"2OBTYOIM","created_at":"2026-07-05T00:59:54Z"}],"graph_snapshots":[{"event_id":"sha256:7256c4f993747898f657645393b220f6aab1d483881d6f381b85b94e2b5156f1","target":"graph","created_at":"2026-07-05T00:59:54Z","signer":{"key_id":"pith-v1-2026-05","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54","signer_id":"pith.science","signer_type":"pith_registry"},"payload":{"graph_snapshot":{"author_claims":{"count":0,"snapshot_sha256":"258153158e38e3291e3d48162225fcdb2d5a3ed65a07baac614ab91432fd4f57","strong_count":0},"builder_version":"pith-number-builder-2026-05-17-v1","claims":{"count":0,"items":[],"snapshot_sha256":"258153158e38e3291e3d48162225fcdb2d5a3ed65a07baac614ab91432fd4f57"},"formal_canon":{"evidence_count":0,"snapshot_sha256":"258153158e38e3291e3d48162225fcdb2d5a3ed65a07baac614ab91432fd4f57"},"integrity":{"available":true,"clean":true,"detectors_run":[],"endpoint":"/pith/1910.14645/integrity.json","findings":[],"snapshot_sha256":"c28c3603d3b5d939e8dc4c7e95fa8dfce3d595e45f758748cecf8e644a296938","summary":{"advisory":0,"by_detector":{},"critical":0,"informational":0}},"paper":{"abstract_excerpt":"During development, organisms acquire three-dimensional shapes with important physiological consequences. While the basic mechanisms underlying morphogenesis are known in eukaryotes, it is often difficult to manipulate them in vivo. To circumvent this issue, here we present a study of developing Vibrio cholerae biofilms grown on agar substrates in which the spatiotemporal morphological patterns were altered by varying the agar concentration. Expanding biofilms are initially flat, but later experience a mechanical instability and become wrinkled. Whereas the peripheral region develops ordered r","authors_text":"Andrej Kosmrlj, Bonnie L. Bassler, Chenyi Fei, Howard A. Stone, Jing Yan, Ned S. Wingreen, Ricard Alert, Sheng Mao","cross_cats":["cond-mat.soft"],"headline":"","license":"http://arxiv.org/licenses/nonexclusive-distrib/1.0/","primary_cat":"physics.bio-ph","submitted_at":"2019-10-31T17:35:00Z","title":"Non-uniform growth and surface friction determine bacterial biofilm morphology on soft substrates"},"references":{"count":0,"internal_anchors":0,"resolved_work":0,"sample":[],"snapshot_sha256":"258153158e38e3291e3d48162225fcdb2d5a3ed65a07baac614ab91432fd4f57"},"source":{"id":"1910.14645","kind":"arxiv","version":1},"verdict":{"created_at":null,"id":null,"model_set":{},"one_line_summary":"","pipeline_version":null,"pith_extraction_headline":"","strongest_claim":"","weakest_assumption":""}},"verdict_id":null}}],"author_attestations":[],"timestamp_anchors":[],"storage_attestations":[],"citation_signatures":[],"replication_records":[],"corrections":[],"mirror_hints":[],"record_created":{"event_id":"sha256:d4e3e9edf6eb23a472adc40e0a88579e4338ce1faa32d842d31339d56e494846","target":"record","created_at":"2026-07-05T00:59:54Z","signer":{"key_id":"pith-v1-2026-05","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54","signer_id":"pith.science","signer_type":"pith_registry"},"payload":{"attestation_state":"computed","canonical_record":{"metadata":{"abstract_canon_sha256":"7accdaa4e3a1f05205aa7131a24d51ed49576e1274967b66bc344238cbf46511","cross_cats_sorted":["cond-mat.soft"],"license":"http://arxiv.org/licenses/nonexclusive-distrib/1.0/","primary_cat":"physics.bio-ph","submitted_at":"2019-10-31T17:35:00Z","title_canon_sha256":"be9a83831eaca612dfca2ff729e8dcc4c70c08a6ec9b60a420dc885e1f77486d"},"schema_version":"1.0","source":{"id":"1910.14645","kind":"arxiv","version":1}},"canonical_sha256":"d3833c390ce4e605ecfc06a4f7260fafc661d04fae1045c3f256d0d3d5e10dcb","receipt":{"algorithm":"ed25519","builder_version":"pith-number-builder-2026-05-17-v1","canonical_sha256":"d3833c390ce4e605ecfc06a4f7260fafc661d04fae1045c3f256d0d3d5e10dcb","first_computed_at":"2026-07-05T00:59:54.972092Z","key_id":"pith-v1-2026-05","kind":"pith_receipt","last_reissued_at":"2026-07-05T00:59:54.972092Z","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54","receipt_version":"0.3","signature_b64":"xSd+rHkTX8M1XOrU6I1+ZMxw+eX6gzewLC8vBMQ6dyB0uynmEtRTWFdF+5TBjWXdqX6ZO7UENN3QHh0RZg9DDQ==","signature_status":"signed_v1","signed_at":"2026-07-05T00:59:54.972626Z","signed_message":"canonical_sha256_bytes"},"source_id":"1910.14645","source_kind":"arxiv","source_version":1}}},"equivocations":[],"invalid_events":[],"applied_event_ids":["sha256:d4e3e9edf6eb23a472adc40e0a88579e4338ce1faa32d842d31339d56e494846","sha256:7256c4f993747898f657645393b220f6aab1d483881d6f381b85b94e2b5156f1"],"state_sha256":"9285ce18c1926cd57f963e3d102975677f7f82c4fb0ef06d8220e372ccacbd95"},"bundle_signature":{"signature_status":"signed_v1","algorithm":"ed25519","key_id":"pith-v1-2026-05","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54","signature_b64":"TAzsaM1il7B6RnTE7ITDy5wlLKrEoY4HX81vmVy86C91HtP4FEpKArh6KHdyzNUV4QaDu5kAU2cpIdnNpA0bAA==","signed_message":"bundle_sha256_bytes","signed_at":"2026-08-04T00:58:32.966474Z","bundle_sha256":"489e052b9a3e2e1005c0dad18c54016a7a06478548b5efe09345b929020c2b2f"}}