{"bundle_type":"pith_open_graph_bundle","bundle_version":"1.0","pith_number":"pith:2015:ATYZQZTMW3O4S52J5V7KXYPROU","short_pith_number":"pith:ATYZQZTM","canonical_record":{"source":{"id":"1505.04471","kind":"arxiv","version":1},"metadata":{"license":"http://arxiv.org/licenses/nonexclusive-distrib/1.0/","primary_cat":"q-bio.PE","submitted_at":"2015-05-17T22:56:36Z","cross_cats_sorted":[],"title_canon_sha256":"c59977158a8449cd3d0d8507ceed52b861fc73733f3141944dd76da5d981572f","abstract_canon_sha256":"f5e9939a6cde314e0d5ba656423ab389dc2c549a0aff289c67431de61227f0c8"},"schema_version":"1.0"},"canonical_sha256":"04f198666cb6ddc97749ed7eabe1f175322f9a984564a814343097f47b5bc154","source":{"kind":"arxiv","id":"1505.04471","version":1},"source_aliases":[{"alias_kind":"arxiv","alias_value":"1505.04471","created_at":"2026-05-18T02:07:25Z"},{"alias_kind":"arxiv_version","alias_value":"1505.04471v1","created_at":"2026-05-18T02:07:25Z"},{"alias_kind":"doi","alias_value":"10.48550/arxiv.1505.04471","created_at":"2026-05-18T02:07:25Z"},{"alias_kind":"pith_short_12","alias_value":"ATYZQZTMW3O4","created_at":"2026-05-18T12:29:10Z"},{"alias_kind":"pith_short_16","alias_value":"ATYZQZTMW3O4S52J","created_at":"2026-05-18T12:29:10Z"},{"alias_kind":"pith_short_8","alias_value":"ATYZQZTM","created_at":"2026-05-18T12:29:10Z"}],"events":[{"event_type":"record_created","subject_pith_number":"pith:2015:ATYZQZTMW3O4S52J5V7KXYPROU","target":"record","payload":{"canonical_record":{"source":{"id":"1505.04471","kind":"arxiv","version":1},"metadata":{"license":"http://arxiv.org/licenses/nonexclusive-distrib/1.0/","primary_cat":"q-bio.PE","submitted_at":"2015-05-17T22:56:36Z","cross_cats_sorted":[],"title_canon_sha256":"c59977158a8449cd3d0d8507ceed52b861fc73733f3141944dd76da5d981572f","abstract_canon_sha256":"f5e9939a6cde314e0d5ba656423ab389dc2c549a0aff289c67431de61227f0c8"},"schema_version":"1.0"},"canonical_sha256":"04f198666cb6ddc97749ed7eabe1f175322f9a984564a814343097f47b5bc154","receipt":{"kind":"pith_receipt","key_id":"pith-v1-2026-05","algorithm":"ed25519","signed_at":"2026-05-18T02:07:25.368042Z","signature_b64":"hpvAbT90rh6Gxfvg/sm2nJ3T708pYlUwUQt75yFwPcbxVHzqLTgkATF3WXElSeAFIv3vbFN7JkSJpKSgu2WpBg==","signed_message":"canonical_sha256_bytes","builder_version":"pith-number-builder-2026-05-17-v1","receipt_version":"0.3","canonical_sha256":"04f198666cb6ddc97749ed7eabe1f175322f9a984564a814343097f47b5bc154","last_reissued_at":"2026-05-18T02:07:25.367537Z","signature_status":"signed_v1","first_computed_at":"2026-05-18T02:07:25.367537Z","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54"},"source_kind":"arxiv","source_id":"1505.04471","source_version":1,"attestation_state":"computed"},"signer":{"signer_id":"pith.science","signer_type":"pith_registry","key_id":"pith-v1-2026-05","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54"},"created_at":"2026-05-18T02:07:25Z","supersedes":[],"prev_event":null,"signature":{"signature_status":"signed_v1","algorithm":"ed25519","key_id":"pith-v1-2026-05","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54","signature_b64":"HSfAgWga8D0yENijePxWILgg84Z4q50WcCiz9ZAb8k6P7l9eWEnD2/BC1uNwFVYaYbK25xOKLQaZOkJzHw4/Bw==","signed_message":"open_graph_event_sha256_bytes","signed_at":"2026-05-30T10:41:09.826498Z"},"content_sha256":"4b56b24fcafcb35f54066dd8f7f0ccd9ab11fac051ac329cc13a7484fef5b624","schema_version":"1.0","event_id":"sha256:4b56b24fcafcb35f54066dd8f7f0ccd9ab11fac051ac329cc13a7484fef5b624"},{"event_type":"graph_snapshot","subject_pith_number":"pith:2015:ATYZQZTMW3O4S52J5V7KXYPROU","target":"graph","payload":{"graph_snapshot":{"paper":{"title":"Evolutionary Fitness in Variable Environments","license":"http://arxiv.org/licenses/nonexclusive-distrib/1.0/","headline":"","cross_cats":[],"primary_cat":"q-bio.PE","authors_text":"Anna Melbinger, Massimo Vergassola","submitted_at":"2015-05-17T22:56:36Z","abstract_excerpt":"One essential ingredient of evolutionary theory is the concept of fitness as a measure for a species' success in its living conditions. Here, we quantify the effect of environmental fluctuations onto fitness by analytical calculations on a general evolutionary model and by studying corresponding individual-based microscopic models. We demonstrate that not only larger growth rates and viabilities, but also reduced sensitivity to environmental variability substantially increases the fitness. Even for neutral evolution, variability in the growth rates plays the crucial role of strongly reducing t"},"claims":{"count":0,"items":[],"snapshot_sha256":"258153158e38e3291e3d48162225fcdb2d5a3ed65a07baac614ab91432fd4f57"},"source":{"id":"1505.04471","kind":"arxiv","version":1},"verdict":{"id":null,"model_set":{},"created_at":null,"strongest_claim":"","one_line_summary":"","pipeline_version":null,"weakest_assumption":"","pith_extraction_headline":""},"references":{"count":0,"sample":[],"resolved_work":0,"snapshot_sha256":"258153158e38e3291e3d48162225fcdb2d5a3ed65a07baac614ab91432fd4f57","internal_anchors":0},"formal_canon":{"evidence_count":0,"snapshot_sha256":"258153158e38e3291e3d48162225fcdb2d5a3ed65a07baac614ab91432fd4f57"},"author_claims":{"count":0,"strong_count":0,"snapshot_sha256":"258153158e38e3291e3d48162225fcdb2d5a3ed65a07baac614ab91432fd4f57"},"builder_version":"pith-number-builder-2026-05-17-v1"},"verdict_id":null},"signer":{"signer_id":"pith.science","signer_type":"pith_registry","key_id":"pith-v1-2026-05","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54"},"created_at":"2026-05-18T02:07:25Z","supersedes":[],"prev_event":null,"signature":{"signature_status":"signed_v1","algorithm":"ed25519","key_id":"pith-v1-2026-05","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54","signature_b64":"9NgG2mqyR7yzAZmkwj1Gg9WBaYQ5lm6sd1UPAGQGTnSsF6VyOlPihrSY9HpkJPYLdKkKlYWe4VE4ZUe5coIwAg==","signed_message":"open_graph_event_sha256_bytes","signed_at":"2026-05-30T10:41:09.826894Z"},"content_sha256":"5ada3868d751bbf07aec288170984e3fb9763d3a918f04f11c4cf623a1c79d30","schema_version":"1.0","event_id":"sha256:5ada3868d751bbf07aec288170984e3fb9763d3a918f04f11c4cf623a1c79d30"}],"timestamp_proofs":[],"mirror_hints":[{"mirror_type":"https","name":"Pith Resolver","base_url":"https://pith.science","bundle_url":"https://pith.science/pith/ATYZQZTMW3O4S52J5V7KXYPROU/bundle.json","state_url":"https://pith.science/pith/ATYZQZTMW3O4S52J5V7KXYPROU/state.json","well_known_bundle_url":"https://pith.science/.well-known/pith/ATYZQZTMW3O4S52J5V7KXYPROU/bundle.json","status":"primary"}],"public_keys":[{"key_id":"pith-v1-2026-05","algorithm":"ed25519","format":"raw","public_key_b64":"stVStoiQhXFxp4s2pdzPNoqVNBMojDU/fJ2db5S3CbM=","public_key_hex":"b2d552b68890857171a78b36a5dccf368a953413288c353f7c9d9d6f94b709b3","fingerprint_sha256_b32_first128bits":"RVFV5Z2OI2J3ZUO7ERDEBCYNKS","fingerprint_sha256_hex":"8d4b5ee74e4693bcd1df2446408b0d54","rotates_at":null,"url":"https://pith.science/pith-signing-key.json","notes":"Pith uses this Ed25519 key to sign canonical record SHA-256 digests. Verify with: ed25519_verify(public_key, message=canonical_sha256_bytes, signature=base64decode(signature_b64))."}],"merge_version":"pith-open-graph-merge-v1","built_at":"2026-05-30T10:41:09Z","links":{"resolver":"https://pith.science/pith/ATYZQZTMW3O4S52J5V7KXYPROU","bundle":"https://pith.science/pith/ATYZQZTMW3O4S52J5V7KXYPROU/bundle.json","state":"https://pith.science/pith/ATYZQZTMW3O4S52J5V7KXYPROU/state.json","well_known_bundle":"https://pith.science/.well-known/pith/ATYZQZTMW3O4S52J5V7KXYPROU/bundle.json"},"state":{"state_type":"pith_open_graph_state","state_version":"1.0","pith_number":"pith:2015:ATYZQZTMW3O4S52J5V7KXYPROU","merge_version":"pith-open-graph-merge-v1","event_count":2,"valid_event_count":2,"invalid_event_count":0,"equivocation_count":0,"current":{"canonical_record":{"metadata":{"abstract_canon_sha256":"f5e9939a6cde314e0d5ba656423ab389dc2c549a0aff289c67431de61227f0c8","cross_cats_sorted":[],"license":"http://arxiv.org/licenses/nonexclusive-distrib/1.0/","primary_cat":"q-bio.PE","submitted_at":"2015-05-17T22:56:36Z","title_canon_sha256":"c59977158a8449cd3d0d8507ceed52b861fc73733f3141944dd76da5d981572f"},"schema_version":"1.0","source":{"id":"1505.04471","kind":"arxiv","version":1}},"source_aliases":[{"alias_kind":"arxiv","alias_value":"1505.04471","created_at":"2026-05-18T02:07:25Z"},{"alias_kind":"arxiv_version","alias_value":"1505.04471v1","created_at":"2026-05-18T02:07:25Z"},{"alias_kind":"doi","alias_value":"10.48550/arxiv.1505.04471","created_at":"2026-05-18T02:07:25Z"},{"alias_kind":"pith_short_12","alias_value":"ATYZQZTMW3O4","created_at":"2026-05-18T12:29:10Z"},{"alias_kind":"pith_short_16","alias_value":"ATYZQZTMW3O4S52J","created_at":"2026-05-18T12:29:10Z"},{"alias_kind":"pith_short_8","alias_value":"ATYZQZTM","created_at":"2026-05-18T12:29:10Z"}],"graph_snapshots":[{"event_id":"sha256:5ada3868d751bbf07aec288170984e3fb9763d3a918f04f11c4cf623a1c79d30","target":"graph","created_at":"2026-05-18T02:07:25Z","signer":{"key_id":"pith-v1-2026-05","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54","signer_id":"pith.science","signer_type":"pith_registry"},"payload":{"graph_snapshot":{"author_claims":{"count":0,"snapshot_sha256":"258153158e38e3291e3d48162225fcdb2d5a3ed65a07baac614ab91432fd4f57","strong_count":0},"builder_version":"pith-number-builder-2026-05-17-v1","claims":{"count":0,"items":[],"snapshot_sha256":"258153158e38e3291e3d48162225fcdb2d5a3ed65a07baac614ab91432fd4f57"},"formal_canon":{"evidence_count":0,"snapshot_sha256":"258153158e38e3291e3d48162225fcdb2d5a3ed65a07baac614ab91432fd4f57"},"paper":{"abstract_excerpt":"One essential ingredient of evolutionary theory is the concept of fitness as a measure for a species' success in its living conditions. Here, we quantify the effect of environmental fluctuations onto fitness by analytical calculations on a general evolutionary model and by studying corresponding individual-based microscopic models. We demonstrate that not only larger growth rates and viabilities, but also reduced sensitivity to environmental variability substantially increases the fitness. Even for neutral evolution, variability in the growth rates plays the crucial role of strongly reducing t","authors_text":"Anna Melbinger, Massimo Vergassola","cross_cats":[],"headline":"","license":"http://arxiv.org/licenses/nonexclusive-distrib/1.0/","primary_cat":"q-bio.PE","submitted_at":"2015-05-17T22:56:36Z","title":"Evolutionary Fitness in Variable Environments"},"references":{"count":0,"internal_anchors":0,"resolved_work":0,"sample":[],"snapshot_sha256":"258153158e38e3291e3d48162225fcdb2d5a3ed65a07baac614ab91432fd4f57"},"source":{"id":"1505.04471","kind":"arxiv","version":1},"verdict":{"created_at":null,"id":null,"model_set":{},"one_line_summary":"","pipeline_version":null,"pith_extraction_headline":"","strongest_claim":"","weakest_assumption":""}},"verdict_id":null}}],"author_attestations":[],"timestamp_anchors":[],"storage_attestations":[],"citation_signatures":[],"replication_records":[],"corrections":[],"mirror_hints":[],"record_created":{"event_id":"sha256:4b56b24fcafcb35f54066dd8f7f0ccd9ab11fac051ac329cc13a7484fef5b624","target":"record","created_at":"2026-05-18T02:07:25Z","signer":{"key_id":"pith-v1-2026-05","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54","signer_id":"pith.science","signer_type":"pith_registry"},"payload":{"attestation_state":"computed","canonical_record":{"metadata":{"abstract_canon_sha256":"f5e9939a6cde314e0d5ba656423ab389dc2c549a0aff289c67431de61227f0c8","cross_cats_sorted":[],"license":"http://arxiv.org/licenses/nonexclusive-distrib/1.0/","primary_cat":"q-bio.PE","submitted_at":"2015-05-17T22:56:36Z","title_canon_sha256":"c59977158a8449cd3d0d8507ceed52b861fc73733f3141944dd76da5d981572f"},"schema_version":"1.0","source":{"id":"1505.04471","kind":"arxiv","version":1}},"canonical_sha256":"04f198666cb6ddc97749ed7eabe1f175322f9a984564a814343097f47b5bc154","receipt":{"algorithm":"ed25519","builder_version":"pith-number-builder-2026-05-17-v1","canonical_sha256":"04f198666cb6ddc97749ed7eabe1f175322f9a984564a814343097f47b5bc154","first_computed_at":"2026-05-18T02:07:25.367537Z","key_id":"pith-v1-2026-05","kind":"pith_receipt","last_reissued_at":"2026-05-18T02:07:25.367537Z","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54","receipt_version":"0.3","signature_b64":"hpvAbT90rh6Gxfvg/sm2nJ3T708pYlUwUQt75yFwPcbxVHzqLTgkATF3WXElSeAFIv3vbFN7JkSJpKSgu2WpBg==","signature_status":"signed_v1","signed_at":"2026-05-18T02:07:25.368042Z","signed_message":"canonical_sha256_bytes"},"source_id":"1505.04471","source_kind":"arxiv","source_version":1}}},"equivocations":[],"invalid_events":[],"applied_event_ids":["sha256:4b56b24fcafcb35f54066dd8f7f0ccd9ab11fac051ac329cc13a7484fef5b624","sha256:5ada3868d751bbf07aec288170984e3fb9763d3a918f04f11c4cf623a1c79d30"],"state_sha256":"40aeb4db9af6d29dd56a3e57bcd67175bc74908a02df02721b1bf94d0753aad1"},"bundle_signature":{"signature_status":"signed_v1","algorithm":"ed25519","key_id":"pith-v1-2026-05","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54","signature_b64":"cuCo3WaYpBtEjcVOpRKZWIe9NQhmeCHlb8DT1qQvf/KBwDp89p5BNBJwomQD7Qf3Mvo3mjOX/tj9ZJZxsWbfBA==","signed_message":"bundle_sha256_bytes","signed_at":"2026-05-30T10:41:09.829494Z","bundle_sha256":"06eae98f6a8ae8ae54e004644690e62d6e722c92ac56b643123a7115d1b41212"}}