{"bundle_type":"pith_open_graph_bundle","bundle_version":"1.0","pith_number":"pith:2011:FVO5NECB5U6FBMUT6KGN5SCREJ","short_pith_number":"pith:FVO5NECB","canonical_record":{"source":{"id":"1112.5804","kind":"arxiv","version":4},"metadata":{"license":"http://arxiv.org/licenses/nonexclusive-distrib/1.0/","primary_cat":"cond-mat.soft","submitted_at":"2011-12-26T02:05:21Z","cross_cats_sorted":["q-bio.CB"],"title_canon_sha256":"c4afcc98918d0bcc37083caee5733417d99e04ceb1ba8818c5f652b09761acaf","abstract_canon_sha256":"dba3b13e0212704574365c9d54a4023e40eca84e7029eed08eea5629826b6cea"},"schema_version":"1.0"},"canonical_sha256":"2d5dd69041ed3c50b293f28cdec8512255df3b3e47ae51971a864defc490a562","source":{"kind":"arxiv","id":"1112.5804","version":4},"source_aliases":[{"alias_kind":"arxiv","alias_value":"1112.5804","created_at":"2026-05-18T01:59:00Z"},{"alias_kind":"arxiv_version","alias_value":"1112.5804v4","created_at":"2026-05-18T01:59:00Z"},{"alias_kind":"doi","alias_value":"10.48550/arxiv.1112.5804","created_at":"2026-05-18T01:59:00Z"},{"alias_kind":"pith_short_12","alias_value":"FVO5NECB5U6F","created_at":"2026-05-18T12:26:28Z"},{"alias_kind":"pith_short_16","alias_value":"FVO5NECB5U6FBMUT","created_at":"2026-05-18T12:26:28Z"},{"alias_kind":"pith_short_8","alias_value":"FVO5NECB","created_at":"2026-05-18T12:26:28Z"}],"events":[{"event_type":"record_created","subject_pith_number":"pith:2011:FVO5NECB5U6FBMUT6KGN5SCREJ","target":"record","payload":{"canonical_record":{"source":{"id":"1112.5804","kind":"arxiv","version":4},"metadata":{"license":"http://arxiv.org/licenses/nonexclusive-distrib/1.0/","primary_cat":"cond-mat.soft","submitted_at":"2011-12-26T02:05:21Z","cross_cats_sorted":["q-bio.CB"],"title_canon_sha256":"c4afcc98918d0bcc37083caee5733417d99e04ceb1ba8818c5f652b09761acaf","abstract_canon_sha256":"dba3b13e0212704574365c9d54a4023e40eca84e7029eed08eea5629826b6cea"},"schema_version":"1.0"},"canonical_sha256":"2d5dd69041ed3c50b293f28cdec8512255df3b3e47ae51971a864defc490a562","receipt":{"kind":"pith_receipt","key_id":"pith-v1-2026-05","algorithm":"ed25519","signed_at":"2026-05-18T01:59:00.649975Z","signature_b64":"U06rKDx2rGTDT2g4MSsLhzjbYJxANL3C90Y82XSLxw3h7SiEhWBk7oFELKRS7h7EAw0PwQKYrTwn/CJLoY54Dg==","signed_message":"canonical_sha256_bytes","builder_version":"pith-number-builder-2026-05-17-v1","receipt_version":"0.3","canonical_sha256":"2d5dd69041ed3c50b293f28cdec8512255df3b3e47ae51971a864defc490a562","last_reissued_at":"2026-05-18T01:59:00.649290Z","signature_status":"signed_v1","first_computed_at":"2026-05-18T01:59:00.649290Z","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54"},"source_kind":"arxiv","source_id":"1112.5804","source_version":4,"attestation_state":"computed"},"signer":{"signer_id":"pith.science","signer_type":"pith_registry","key_id":"pith-v1-2026-05","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54"},"created_at":"2026-05-18T01:59:00Z","supersedes":[],"prev_event":null,"signature":{"signature_status":"signed_v1","algorithm":"ed25519","key_id":"pith-v1-2026-05","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54","signature_b64":"cMKpXiA1yoJYldom6sYU6lo/O9Fq3JwbU7eNexo+dVQYLDxX2/hFIlzqEEbxd/QRw3ARp/A2GqU2qi2Kibo3DA==","signed_message":"open_graph_event_sha256_bytes","signed_at":"2026-05-27T03:31:40.713941Z"},"content_sha256":"090e7d4ea994bec9ceb31ea9d5c09ab1181fa5da1494e604338b56950016d996","schema_version":"1.0","event_id":"sha256:090e7d4ea994bec9ceb31ea9d5c09ab1181fa5da1494e604338b56950016d996"},{"event_type":"graph_snapshot","subject_pith_number":"pith:2011:FVO5NECB5U6FBMUT6KGN5SCREJ","target":"graph","payload":{"graph_snapshot":{"paper":{"title":"Scaling of Traction Forces with Size of Cohesive Cell Colonies","license":"http://arxiv.org/licenses/nonexclusive-distrib/1.0/","headline":"","cross_cats":["q-bio.CB"],"primary_cat":"cond-mat.soft","authors_text":"Aaron F. Mertz, Callen Hyland, Eric. R. Dufresne, Guy K. German, M. Cristina Marchetti, Shiladitya Banerjee, Valerie Horsley, Ye Xu, Yonglu Che","submitted_at":"2011-12-26T02:05:21Z","abstract_excerpt":"To understand how the mechanical properties of tissues emerge from interactions of multiple cells, we measure traction stresses of cohesive colonies of 1-27 cells adherent to soft substrates. We find that traction stresses are generally localized at the periphery of the colony and the total traction force scales with the colony radius. For large colony sizes, the scaling appears to approach linear, suggesting the emergence of an apparent surface tension of order 1E-3 N/m. A simple model of the cell colony as a contractile elastic medium coupled to the substrate captures the spatial distributio"},"claims":{"count":0,"items":[],"snapshot_sha256":"258153158e38e3291e3d48162225fcdb2d5a3ed65a07baac614ab91432fd4f57"},"source":{"id":"1112.5804","kind":"arxiv","version":4},"verdict":{"id":null,"model_set":{},"created_at":null,"strongest_claim":"","one_line_summary":"","pipeline_version":null,"weakest_assumption":"","pith_extraction_headline":""},"references":{"count":0,"sample":[],"resolved_work":0,"snapshot_sha256":"258153158e38e3291e3d48162225fcdb2d5a3ed65a07baac614ab91432fd4f57","internal_anchors":0},"formal_canon":{"evidence_count":0,"snapshot_sha256":"258153158e38e3291e3d48162225fcdb2d5a3ed65a07baac614ab91432fd4f57"},"author_claims":{"count":0,"strong_count":0,"snapshot_sha256":"258153158e38e3291e3d48162225fcdb2d5a3ed65a07baac614ab91432fd4f57"},"builder_version":"pith-number-builder-2026-05-17-v1"},"verdict_id":null},"signer":{"signer_id":"pith.science","signer_type":"pith_registry","key_id":"pith-v1-2026-05","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54"},"created_at":"2026-05-18T01:59:00Z","supersedes":[],"prev_event":null,"signature":{"signature_status":"signed_v1","algorithm":"ed25519","key_id":"pith-v1-2026-05","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54","signature_b64":"8C8DWk55qx3QDXKoDdDhQ+YFHOFUJiYg2A3/iIcveoWuCQZU91osvxKI7VZ3rcgJeYjqT5I1Wn+FHDuHgqNYBw==","signed_message":"open_graph_event_sha256_bytes","signed_at":"2026-05-27T03:31:40.714621Z"},"content_sha256":"a283733ee5f38ae4060b0a331d3b925839709f9ed67b4133bca45c6b08edd962","schema_version":"1.0","event_id":"sha256:a283733ee5f38ae4060b0a331d3b925839709f9ed67b4133bca45c6b08edd962"}],"timestamp_proofs":[],"mirror_hints":[{"mirror_type":"https","name":"Pith Resolver","base_url":"https://pith.science","bundle_url":"https://pith.science/pith/FVO5NECB5U6FBMUT6KGN5SCREJ/bundle.json","state_url":"https://pith.science/pith/FVO5NECB5U6FBMUT6KGN5SCREJ/state.json","well_known_bundle_url":"https://pith.science/.well-known/pith/FVO5NECB5U6FBMUT6KGN5SCREJ/bundle.json","status":"primary"}],"public_keys":[{"key_id":"pith-v1-2026-05","algorithm":"ed25519","format":"raw","public_key_b64":"stVStoiQhXFxp4s2pdzPNoqVNBMojDU/fJ2db5S3CbM=","public_key_hex":"b2d552b68890857171a78b36a5dccf368a953413288c353f7c9d9d6f94b709b3","fingerprint_sha256_b32_first128bits":"RVFV5Z2OI2J3ZUO7ERDEBCYNKS","fingerprint_sha256_hex":"8d4b5ee74e4693bcd1df2446408b0d54","rotates_at":null,"url":"https://pith.science/pith-signing-key.json","notes":"Pith uses this Ed25519 key to sign canonical record SHA-256 digests. Verify with: ed25519_verify(public_key, message=canonical_sha256_bytes, signature=base64decode(signature_b64))."}],"merge_version":"pith-open-graph-merge-v1","built_at":"2026-05-27T03:31:40Z","links":{"resolver":"https://pith.science/pith/FVO5NECB5U6FBMUT6KGN5SCREJ","bundle":"https://pith.science/pith/FVO5NECB5U6FBMUT6KGN5SCREJ/bundle.json","state":"https://pith.science/pith/FVO5NECB5U6FBMUT6KGN5SCREJ/state.json","well_known_bundle":"https://pith.science/.well-known/pith/FVO5NECB5U6FBMUT6KGN5SCREJ/bundle.json"},"state":{"state_type":"pith_open_graph_state","state_version":"1.0","pith_number":"pith:2011:FVO5NECB5U6FBMUT6KGN5SCREJ","merge_version":"pith-open-graph-merge-v1","event_count":2,"valid_event_count":2,"invalid_event_count":0,"equivocation_count":0,"current":{"canonical_record":{"metadata":{"abstract_canon_sha256":"dba3b13e0212704574365c9d54a4023e40eca84e7029eed08eea5629826b6cea","cross_cats_sorted":["q-bio.CB"],"license":"http://arxiv.org/licenses/nonexclusive-distrib/1.0/","primary_cat":"cond-mat.soft","submitted_at":"2011-12-26T02:05:21Z","title_canon_sha256":"c4afcc98918d0bcc37083caee5733417d99e04ceb1ba8818c5f652b09761acaf"},"schema_version":"1.0","source":{"id":"1112.5804","kind":"arxiv","version":4}},"source_aliases":[{"alias_kind":"arxiv","alias_value":"1112.5804","created_at":"2026-05-18T01:59:00Z"},{"alias_kind":"arxiv_version","alias_value":"1112.5804v4","created_at":"2026-05-18T01:59:00Z"},{"alias_kind":"doi","alias_value":"10.48550/arxiv.1112.5804","created_at":"2026-05-18T01:59:00Z"},{"alias_kind":"pith_short_12","alias_value":"FVO5NECB5U6F","created_at":"2026-05-18T12:26:28Z"},{"alias_kind":"pith_short_16","alias_value":"FVO5NECB5U6FBMUT","created_at":"2026-05-18T12:26:28Z"},{"alias_kind":"pith_short_8","alias_value":"FVO5NECB","created_at":"2026-05-18T12:26:28Z"}],"graph_snapshots":[{"event_id":"sha256:a283733ee5f38ae4060b0a331d3b925839709f9ed67b4133bca45c6b08edd962","target":"graph","created_at":"2026-05-18T01:59:00Z","signer":{"key_id":"pith-v1-2026-05","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54","signer_id":"pith.science","signer_type":"pith_registry"},"payload":{"graph_snapshot":{"author_claims":{"count":0,"snapshot_sha256":"258153158e38e3291e3d48162225fcdb2d5a3ed65a07baac614ab91432fd4f57","strong_count":0},"builder_version":"pith-number-builder-2026-05-17-v1","claims":{"count":0,"items":[],"snapshot_sha256":"258153158e38e3291e3d48162225fcdb2d5a3ed65a07baac614ab91432fd4f57"},"formal_canon":{"evidence_count":0,"snapshot_sha256":"258153158e38e3291e3d48162225fcdb2d5a3ed65a07baac614ab91432fd4f57"},"paper":{"abstract_excerpt":"To understand how the mechanical properties of tissues emerge from interactions of multiple cells, we measure traction stresses of cohesive colonies of 1-27 cells adherent to soft substrates. We find that traction stresses are generally localized at the periphery of the colony and the total traction force scales with the colony radius. For large colony sizes, the scaling appears to approach linear, suggesting the emergence of an apparent surface tension of order 1E-3 N/m. A simple model of the cell colony as a contractile elastic medium coupled to the substrate captures the spatial distributio","authors_text":"Aaron F. Mertz, Callen Hyland, Eric. R. Dufresne, Guy K. German, M. Cristina Marchetti, Shiladitya Banerjee, Valerie Horsley, Ye Xu, Yonglu Che","cross_cats":["q-bio.CB"],"headline":"","license":"http://arxiv.org/licenses/nonexclusive-distrib/1.0/","primary_cat":"cond-mat.soft","submitted_at":"2011-12-26T02:05:21Z","title":"Scaling of Traction Forces with Size of Cohesive Cell Colonies"},"references":{"count":0,"internal_anchors":0,"resolved_work":0,"sample":[],"snapshot_sha256":"258153158e38e3291e3d48162225fcdb2d5a3ed65a07baac614ab91432fd4f57"},"source":{"id":"1112.5804","kind":"arxiv","version":4},"verdict":{"created_at":null,"id":null,"model_set":{},"one_line_summary":"","pipeline_version":null,"pith_extraction_headline":"","strongest_claim":"","weakest_assumption":""}},"verdict_id":null}}],"author_attestations":[],"timestamp_anchors":[],"storage_attestations":[],"citation_signatures":[],"replication_records":[],"corrections":[],"mirror_hints":[],"record_created":{"event_id":"sha256:090e7d4ea994bec9ceb31ea9d5c09ab1181fa5da1494e604338b56950016d996","target":"record","created_at":"2026-05-18T01:59:00Z","signer":{"key_id":"pith-v1-2026-05","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54","signer_id":"pith.science","signer_type":"pith_registry"},"payload":{"attestation_state":"computed","canonical_record":{"metadata":{"abstract_canon_sha256":"dba3b13e0212704574365c9d54a4023e40eca84e7029eed08eea5629826b6cea","cross_cats_sorted":["q-bio.CB"],"license":"http://arxiv.org/licenses/nonexclusive-distrib/1.0/","primary_cat":"cond-mat.soft","submitted_at":"2011-12-26T02:05:21Z","title_canon_sha256":"c4afcc98918d0bcc37083caee5733417d99e04ceb1ba8818c5f652b09761acaf"},"schema_version":"1.0","source":{"id":"1112.5804","kind":"arxiv","version":4}},"canonical_sha256":"2d5dd69041ed3c50b293f28cdec8512255df3b3e47ae51971a864defc490a562","receipt":{"algorithm":"ed25519","builder_version":"pith-number-builder-2026-05-17-v1","canonical_sha256":"2d5dd69041ed3c50b293f28cdec8512255df3b3e47ae51971a864defc490a562","first_computed_at":"2026-05-18T01:59:00.649290Z","key_id":"pith-v1-2026-05","kind":"pith_receipt","last_reissued_at":"2026-05-18T01:59:00.649290Z","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54","receipt_version":"0.3","signature_b64":"U06rKDx2rGTDT2g4MSsLhzjbYJxANL3C90Y82XSLxw3h7SiEhWBk7oFELKRS7h7EAw0PwQKYrTwn/CJLoY54Dg==","signature_status":"signed_v1","signed_at":"2026-05-18T01:59:00.649975Z","signed_message":"canonical_sha256_bytes"},"source_id":"1112.5804","source_kind":"arxiv","source_version":4}}},"equivocations":[],"invalid_events":[],"applied_event_ids":["sha256:090e7d4ea994bec9ceb31ea9d5c09ab1181fa5da1494e604338b56950016d996","sha256:a283733ee5f38ae4060b0a331d3b925839709f9ed67b4133bca45c6b08edd962"],"state_sha256":"8d2ac9a0086f3332864fbd31ce08ff755fde1b23a580b1816291351646d4b6ba"},"bundle_signature":{"signature_status":"signed_v1","algorithm":"ed25519","key_id":"pith-v1-2026-05","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54","signature_b64":"8Iim+op+sBtZVtVCuXFaOFCcwEi8aqy8+9Bm8KlWOfAVzmYPW1bZLhwClOMLL5FI1/5AAlqigHgID6xczUs+Aw==","signed_message":"bundle_sha256_bytes","signed_at":"2026-05-27T03:31:40.718063Z","bundle_sha256":"336eda5def5a2ac21037c36d81d3e1aa77df96e5e83c229daced75008e05bfee"}}