{"bundle_type":"pith_open_graph_bundle","bundle_version":"1.0","pith_number":"pith:2013:MFS3NY2IQEIUIJMBA4YUYQHGJD","short_pith_number":"pith:MFS3NY2I","canonical_record":{"source":{"id":"1303.5044","kind":"arxiv","version":3},"metadata":{"license":"http://arxiv.org/licenses/nonexclusive-distrib/1.0/","primary_cat":"q-bio.PE","submitted_at":"2013-03-20T19:42:07Z","cross_cats_sorted":[],"title_canon_sha256":"cf9642a4a14fdffea869c875ac7267d5547d09b480cb466696b8dad5686e2c03","abstract_canon_sha256":"02d09b1bbb90bd09c782c690c6c33bdea8a0fd2a27b4e0e2a4d239952ba0bac4"},"schema_version":"1.0"},"canonical_sha256":"6165b6e348811144258107314c40e648fba4db687b0f1913fb18768c2e4a714c","source":{"kind":"arxiv","id":"1303.5044","version":3},"source_aliases":[{"alias_kind":"arxiv","alias_value":"1303.5044","created_at":"2026-05-18T02:38:26Z"},{"alias_kind":"arxiv_version","alias_value":"1303.5044v3","created_at":"2026-05-18T02:38:26Z"},{"alias_kind":"doi","alias_value":"10.48550/arxiv.1303.5044","created_at":"2026-05-18T02:38:26Z"},{"alias_kind":"pith_short_12","alias_value":"MFS3NY2IQEIU","created_at":"2026-05-18T12:27:52Z"},{"alias_kind":"pith_short_16","alias_value":"MFS3NY2IQEIUIJMB","created_at":"2026-05-18T12:27:52Z"},{"alias_kind":"pith_short_8","alias_value":"MFS3NY2I","created_at":"2026-05-18T12:27:52Z"}],"events":[{"event_type":"record_created","subject_pith_number":"pith:2013:MFS3NY2IQEIUIJMBA4YUYQHGJD","target":"record","payload":{"canonical_record":{"source":{"id":"1303.5044","kind":"arxiv","version":3},"metadata":{"license":"http://arxiv.org/licenses/nonexclusive-distrib/1.0/","primary_cat":"q-bio.PE","submitted_at":"2013-03-20T19:42:07Z","cross_cats_sorted":[],"title_canon_sha256":"cf9642a4a14fdffea869c875ac7267d5547d09b480cb466696b8dad5686e2c03","abstract_canon_sha256":"02d09b1bbb90bd09c782c690c6c33bdea8a0fd2a27b4e0e2a4d239952ba0bac4"},"schema_version":"1.0"},"canonical_sha256":"6165b6e348811144258107314c40e648fba4db687b0f1913fb18768c2e4a714c","receipt":{"kind":"pith_receipt","key_id":"pith-v1-2026-05","algorithm":"ed25519","signed_at":"2026-05-18T02:38:26.740340Z","signature_b64":"sZ6htcOiv/+jc934gXJDUVxIVCcedk5v9EDioSPOSTLqH9pGeZFEL7mURhl4RaQLGkdyn69mYXeEUOp5DMasCg==","signed_message":"canonical_sha256_bytes","builder_version":"pith-number-builder-2026-05-17-v1","receipt_version":"0.3","canonical_sha256":"6165b6e348811144258107314c40e648fba4db687b0f1913fb18768c2e4a714c","last_reissued_at":"2026-05-18T02:38:26.739682Z","signature_status":"signed_v1","first_computed_at":"2026-05-18T02:38:26.739682Z","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54"},"source_kind":"arxiv","source_id":"1303.5044","source_version":3,"attestation_state":"computed"},"signer":{"signer_id":"pith.science","signer_type":"pith_registry","key_id":"pith-v1-2026-05","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54"},"created_at":"2026-05-18T02:38:26Z","supersedes":[],"prev_event":null,"signature":{"signature_status":"signed_v1","algorithm":"ed25519","key_id":"pith-v1-2026-05","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54","signature_b64":"gujKlkX0NHxGAAKHGYqCbWzHTz12p/7vois0IcFF4iAA68cRk44oSJEfKQf1W7VE7iOnzb8cKTcAfXqz5RpEAQ==","signed_message":"open_graph_event_sha256_bytes","signed_at":"2026-06-30T15:15:15.559428Z"},"content_sha256":"068aee8909f48ded3c11beeae3a0507b39a63e26d97d531d99549c45e7622b52","schema_version":"1.0","event_id":"sha256:068aee8909f48ded3c11beeae3a0507b39a63e26d97d531d99549c45e7622b52"},{"event_type":"graph_snapshot","subject_pith_number":"pith:2013:MFS3NY2IQEIUIJMBA4YUYQHGJD","target":"graph","payload":{"graph_snapshot":{"paper":{"title":"Genomic evidence of rapid and stable adaptive oscillations over seasonal time scales in Drosophila","license":"http://arxiv.org/licenses/nonexclusive-distrib/1.0/","headline":"","cross_cats":[],"primary_cat":"q-bio.PE","authors_text":"Alan O. Bergland, Dmitri A. Petrov, Emily L. Behrman, Katherine R. O'Brien, Paul S. Schmidt","submitted_at":"2013-03-20T19:42:07Z","abstract_excerpt":"In many species, genomic data have revealed pervasive adaptive evolution indicated by the fixation of beneficial alleles. However, when selection pressures are highly variable along a species range or through time adaptive alleles may persist at intermediate frequencies for long periods. So called balanced polymorphisms have long been understood to be an important component of standing genetic variation yet direct evidence of the strength of balancing selection and the stability and prevalence of balanced polymorphisms has remained elusive. We hypothesized that environmental fluctuations betwe"},"claims":{"count":0,"items":[],"snapshot_sha256":"258153158e38e3291e3d48162225fcdb2d5a3ed65a07baac614ab91432fd4f57"},"source":{"id":"1303.5044","kind":"arxiv","version":3},"verdict":{"id":null,"model_set":{},"created_at":null,"strongest_claim":"","one_line_summary":"","pipeline_version":null,"weakest_assumption":"","pith_extraction_headline":""},"references":{"count":0,"sample":[],"resolved_work":0,"snapshot_sha256":"258153158e38e3291e3d48162225fcdb2d5a3ed65a07baac614ab91432fd4f57","internal_anchors":0},"formal_canon":{"evidence_count":0,"snapshot_sha256":"258153158e38e3291e3d48162225fcdb2d5a3ed65a07baac614ab91432fd4f57"},"author_claims":{"count":0,"strong_count":0,"snapshot_sha256":"258153158e38e3291e3d48162225fcdb2d5a3ed65a07baac614ab91432fd4f57"},"builder_version":"pith-number-builder-2026-05-17-v1"},"verdict_id":null},"signer":{"signer_id":"pith.science","signer_type":"pith_registry","key_id":"pith-v1-2026-05","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54"},"created_at":"2026-05-18T02:38:26Z","supersedes":[],"prev_event":null,"signature":{"signature_status":"signed_v1","algorithm":"ed25519","key_id":"pith-v1-2026-05","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54","signature_b64":"W7h20cViuamRjqKPpYBGMtmeRkIJQg2hV4OEzzjdDGa+jLE5wBAYJr6+WVISQnzaNSOY7PyMAxazWTQEFmVECg==","signed_message":"open_graph_event_sha256_bytes","signed_at":"2026-06-30T15:15:15.559785Z"},"content_sha256":"329bd834801a4967183abff492fad7b00c5d48f5762729c02e3a6d6b34eda507","schema_version":"1.0","event_id":"sha256:329bd834801a4967183abff492fad7b00c5d48f5762729c02e3a6d6b34eda507"}],"timestamp_proofs":[],"mirror_hints":[{"mirror_type":"https","name":"Pith Resolver","base_url":"https://pith.science","bundle_url":"https://pith.science/pith/MFS3NY2IQEIUIJMBA4YUYQHGJD/bundle.json","state_url":"https://pith.science/pith/MFS3NY2IQEIUIJMBA4YUYQHGJD/state.json","well_known_bundle_url":"https://pith.science/.well-known/pith/MFS3NY2IQEIUIJMBA4YUYQHGJD/bundle.json","status":"primary"}],"public_keys":[{"key_id":"pith-v1-2026-05","algorithm":"ed25519","format":"raw","public_key_b64":"stVStoiQhXFxp4s2pdzPNoqVNBMojDU/fJ2db5S3CbM=","public_key_hex":"b2d552b68890857171a78b36a5dccf368a953413288c353f7c9d9d6f94b709b3","fingerprint_sha256_b32_first128bits":"RVFV5Z2OI2J3ZUO7ERDEBCYNKS","fingerprint_sha256_hex":"8d4b5ee74e4693bcd1df2446408b0d54","rotates_at":null,"url":"https://pith.science/pith-signing-key.json","notes":"Pith uses this Ed25519 key to sign canonical record SHA-256 digests. Verify with: ed25519_verify(public_key, message=canonical_sha256_bytes, signature=base64decode(signature_b64))."}],"merge_version":"pith-open-graph-merge-v1","built_at":"2026-06-30T15:15:15Z","links":{"resolver":"https://pith.science/pith/MFS3NY2IQEIUIJMBA4YUYQHGJD","bundle":"https://pith.science/pith/MFS3NY2IQEIUIJMBA4YUYQHGJD/bundle.json","state":"https://pith.science/pith/MFS3NY2IQEIUIJMBA4YUYQHGJD/state.json","well_known_bundle":"https://pith.science/.well-known/pith/MFS3NY2IQEIUIJMBA4YUYQHGJD/bundle.json"},"state":{"state_type":"pith_open_graph_state","state_version":"1.0","pith_number":"pith:2013:MFS3NY2IQEIUIJMBA4YUYQHGJD","merge_version":"pith-open-graph-merge-v1","event_count":2,"valid_event_count":2,"invalid_event_count":0,"equivocation_count":0,"current":{"canonical_record":{"metadata":{"abstract_canon_sha256":"02d09b1bbb90bd09c782c690c6c33bdea8a0fd2a27b4e0e2a4d239952ba0bac4","cross_cats_sorted":[],"license":"http://arxiv.org/licenses/nonexclusive-distrib/1.0/","primary_cat":"q-bio.PE","submitted_at":"2013-03-20T19:42:07Z","title_canon_sha256":"cf9642a4a14fdffea869c875ac7267d5547d09b480cb466696b8dad5686e2c03"},"schema_version":"1.0","source":{"id":"1303.5044","kind":"arxiv","version":3}},"source_aliases":[{"alias_kind":"arxiv","alias_value":"1303.5044","created_at":"2026-05-18T02:38:26Z"},{"alias_kind":"arxiv_version","alias_value":"1303.5044v3","created_at":"2026-05-18T02:38:26Z"},{"alias_kind":"doi","alias_value":"10.48550/arxiv.1303.5044","created_at":"2026-05-18T02:38:26Z"},{"alias_kind":"pith_short_12","alias_value":"MFS3NY2IQEIU","created_at":"2026-05-18T12:27:52Z"},{"alias_kind":"pith_short_16","alias_value":"MFS3NY2IQEIUIJMB","created_at":"2026-05-18T12:27:52Z"},{"alias_kind":"pith_short_8","alias_value":"MFS3NY2I","created_at":"2026-05-18T12:27:52Z"}],"graph_snapshots":[{"event_id":"sha256:329bd834801a4967183abff492fad7b00c5d48f5762729c02e3a6d6b34eda507","target":"graph","created_at":"2026-05-18T02:38:26Z","signer":{"key_id":"pith-v1-2026-05","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54","signer_id":"pith.science","signer_type":"pith_registry"},"payload":{"graph_snapshot":{"author_claims":{"count":0,"snapshot_sha256":"258153158e38e3291e3d48162225fcdb2d5a3ed65a07baac614ab91432fd4f57","strong_count":0},"builder_version":"pith-number-builder-2026-05-17-v1","claims":{"count":0,"items":[],"snapshot_sha256":"258153158e38e3291e3d48162225fcdb2d5a3ed65a07baac614ab91432fd4f57"},"formal_canon":{"evidence_count":0,"snapshot_sha256":"258153158e38e3291e3d48162225fcdb2d5a3ed65a07baac614ab91432fd4f57"},"paper":{"abstract_excerpt":"In many species, genomic data have revealed pervasive adaptive evolution indicated by the fixation of beneficial alleles. However, when selection pressures are highly variable along a species range or through time adaptive alleles may persist at intermediate frequencies for long periods. So called balanced polymorphisms have long been understood to be an important component of standing genetic variation yet direct evidence of the strength of balancing selection and the stability and prevalence of balanced polymorphisms has remained elusive. We hypothesized that environmental fluctuations betwe","authors_text":"Alan O. Bergland, Dmitri A. Petrov, Emily L. Behrman, Katherine R. O'Brien, Paul S. Schmidt","cross_cats":[],"headline":"","license":"http://arxiv.org/licenses/nonexclusive-distrib/1.0/","primary_cat":"q-bio.PE","submitted_at":"2013-03-20T19:42:07Z","title":"Genomic evidence of rapid and stable adaptive oscillations over seasonal time scales in Drosophila"},"references":{"count":0,"internal_anchors":0,"resolved_work":0,"sample":[],"snapshot_sha256":"258153158e38e3291e3d48162225fcdb2d5a3ed65a07baac614ab91432fd4f57"},"source":{"id":"1303.5044","kind":"arxiv","version":3},"verdict":{"created_at":null,"id":null,"model_set":{},"one_line_summary":"","pipeline_version":null,"pith_extraction_headline":"","strongest_claim":"","weakest_assumption":""}},"verdict_id":null}}],"author_attestations":[],"timestamp_anchors":[],"storage_attestations":[],"citation_signatures":[],"replication_records":[],"corrections":[],"mirror_hints":[],"record_created":{"event_id":"sha256:068aee8909f48ded3c11beeae3a0507b39a63e26d97d531d99549c45e7622b52","target":"record","created_at":"2026-05-18T02:38:26Z","signer":{"key_id":"pith-v1-2026-05","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54","signer_id":"pith.science","signer_type":"pith_registry"},"payload":{"attestation_state":"computed","canonical_record":{"metadata":{"abstract_canon_sha256":"02d09b1bbb90bd09c782c690c6c33bdea8a0fd2a27b4e0e2a4d239952ba0bac4","cross_cats_sorted":[],"license":"http://arxiv.org/licenses/nonexclusive-distrib/1.0/","primary_cat":"q-bio.PE","submitted_at":"2013-03-20T19:42:07Z","title_canon_sha256":"cf9642a4a14fdffea869c875ac7267d5547d09b480cb466696b8dad5686e2c03"},"schema_version":"1.0","source":{"id":"1303.5044","kind":"arxiv","version":3}},"canonical_sha256":"6165b6e348811144258107314c40e648fba4db687b0f1913fb18768c2e4a714c","receipt":{"algorithm":"ed25519","builder_version":"pith-number-builder-2026-05-17-v1","canonical_sha256":"6165b6e348811144258107314c40e648fba4db687b0f1913fb18768c2e4a714c","first_computed_at":"2026-05-18T02:38:26.739682Z","key_id":"pith-v1-2026-05","kind":"pith_receipt","last_reissued_at":"2026-05-18T02:38:26.739682Z","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54","receipt_version":"0.3","signature_b64":"sZ6htcOiv/+jc934gXJDUVxIVCcedk5v9EDioSPOSTLqH9pGeZFEL7mURhl4RaQLGkdyn69mYXeEUOp5DMasCg==","signature_status":"signed_v1","signed_at":"2026-05-18T02:38:26.740340Z","signed_message":"canonical_sha256_bytes"},"source_id":"1303.5044","source_kind":"arxiv","source_version":3}}},"equivocations":[],"invalid_events":[],"applied_event_ids":["sha256:068aee8909f48ded3c11beeae3a0507b39a63e26d97d531d99549c45e7622b52","sha256:329bd834801a4967183abff492fad7b00c5d48f5762729c02e3a6d6b34eda507"],"state_sha256":"3e0a9dd8f89ff09ebc8fc2dd59eb4a27525895e03f275c7180af866076b6a53d"},"bundle_signature":{"signature_status":"signed_v1","algorithm":"ed25519","key_id":"pith-v1-2026-05","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54","signature_b64":"Gzt30zE2jI9Col+gBKCE0d2BxUHpEkx2efxIS8EP4eePA2cUHbiETEsp5KVZ75ptDBpkF3jav0VD6/vumPpOBA==","signed_message":"bundle_sha256_bytes","signed_at":"2026-06-30T15:15:15.562190Z","bundle_sha256":"e46d3d0f6010df79191a48a0cad5e2073157d5713aff0b03fd50feea60ae60fa"}}