{"bundle_type":"pith_open_graph_bundle","bundle_version":"1.0","pith_number":"pith:2018:TOZC2JYUECSBABB4FSNA7P3MTQ","short_pith_number":"pith:TOZC2JYU","canonical_record":{"source":{"id":"1807.08719","kind":"arxiv","version":1},"metadata":{"license":"http://arxiv.org/licenses/nonexclusive-distrib/1.0/","primary_cat":"physics.bio-ph","submitted_at":"2018-07-23T17:00:57Z","cross_cats_sorted":["q-bio.BM","q-bio.QM","q-bio.SC"],"title_canon_sha256":"1f520c67e85b1be981e4050352efc001a39271e1db33e603b88daed4b217dc79","abstract_canon_sha256":"3d4e56f7ddeb18472bc7593302363b75564b4d84f53a79440e6a8287b104d165"},"schema_version":"1.0"},"canonical_sha256":"9bb22d271420a410043c2c9a0fbf6c9c0f28d21aee70b41c2ea6d95c2ea9e2ae","source":{"kind":"arxiv","id":"1807.08719","version":1},"source_aliases":[{"alias_kind":"arxiv","alias_value":"1807.08719","created_at":"2026-05-18T00:10:04Z"},{"alias_kind":"arxiv_version","alias_value":"1807.08719v1","created_at":"2026-05-18T00:10:04Z"},{"alias_kind":"doi","alias_value":"10.48550/arxiv.1807.08719","created_at":"2026-05-18T00:10:04Z"},{"alias_kind":"pith_short_12","alias_value":"TOZC2JYUECSB","created_at":"2026-05-18T12:32:53Z"},{"alias_kind":"pith_short_16","alias_value":"TOZC2JYUECSBABB4","created_at":"2026-05-18T12:32:53Z"},{"alias_kind":"pith_short_8","alias_value":"TOZC2JYU","created_at":"2026-05-18T12:32:53Z"}],"events":[{"event_type":"record_created","subject_pith_number":"pith:2018:TOZC2JYUECSBABB4FSNA7P3MTQ","target":"record","payload":{"canonical_record":{"source":{"id":"1807.08719","kind":"arxiv","version":1},"metadata":{"license":"http://arxiv.org/licenses/nonexclusive-distrib/1.0/","primary_cat":"physics.bio-ph","submitted_at":"2018-07-23T17:00:57Z","cross_cats_sorted":["q-bio.BM","q-bio.QM","q-bio.SC"],"title_canon_sha256":"1f520c67e85b1be981e4050352efc001a39271e1db33e603b88daed4b217dc79","abstract_canon_sha256":"3d4e56f7ddeb18472bc7593302363b75564b4d84f53a79440e6a8287b104d165"},"schema_version":"1.0"},"canonical_sha256":"9bb22d271420a410043c2c9a0fbf6c9c0f28d21aee70b41c2ea6d95c2ea9e2ae","receipt":{"kind":"pith_receipt","key_id":"pith-v1-2026-05","algorithm":"ed25519","signed_at":"2026-05-18T00:10:04.898213Z","signature_b64":"wyQKi+UgxRgE5WdPg0P5q5nK8ygo1P9y1O6CbrNy/AO6alXBlsDYqA7qR7pFMXb1Rl7VbUd/ZDQgKq7qr0cMDg==","signed_message":"canonical_sha256_bytes","builder_version":"pith-number-builder-2026-05-17-v1","receipt_version":"0.3","canonical_sha256":"9bb22d271420a410043c2c9a0fbf6c9c0f28d21aee70b41c2ea6d95c2ea9e2ae","last_reissued_at":"2026-05-18T00:10:04.897537Z","signature_status":"signed_v1","first_computed_at":"2026-05-18T00:10:04.897537Z","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54"},"source_kind":"arxiv","source_id":"1807.08719","source_version":1,"attestation_state":"computed"},"signer":{"signer_id":"pith.science","signer_type":"pith_registry","key_id":"pith-v1-2026-05","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54"},"created_at":"2026-05-18T00:10:04Z","supersedes":[],"prev_event":null,"signature":{"signature_status":"signed_v1","algorithm":"ed25519","key_id":"pith-v1-2026-05","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54","signature_b64":"jzuC5Js9StcKPUqQ0C8Tt7/d7BIxGKUX4/79lgHjPJ1NrYGKqJfofCuWPr+odqgyuhXXh0rx9L0TAl316jjyCg==","signed_message":"open_graph_event_sha256_bytes","signed_at":"2026-06-02T14:16:58.868788Z"},"content_sha256":"6ea04fa9a163482f4226b25790c548637e2bdcfed23c6c25d7a74476cf9ca8d3","schema_version":"1.0","event_id":"sha256:6ea04fa9a163482f4226b25790c548637e2bdcfed23c6c25d7a74476cf9ca8d3"},{"event_type":"graph_snapshot","subject_pith_number":"pith:2018:TOZC2JYUECSBABB4FSNA7P3MTQ","target":"graph","payload":{"graph_snapshot":{"paper":{"title":"Microtubules soften due to cross-sectional flattening","license":"http://arxiv.org/licenses/nonexclusive-distrib/1.0/","headline":"","cross_cats":["q-bio.BM","q-bio.QM","q-bio.SC"],"primary_cat":"physics.bio-ph","authors_text":"Edvin Memet, Feodor Hilitski, L Mahadevan, Margaret A Morris, Walter J Schwenger, Zvonimir Dogic","submitted_at":"2018-07-23T17:00:57Z","abstract_excerpt":"We use optical trapping to continuously bend an isolated microtubule while simultaneously measuring the applied force and the resulting filament strain, thus allowing us to determine its elastic properties over a wide range of applied strains. We find that, while in the low-strain regime, microtubules may be quantitatively described in terms of the classical Euler-Bernoulli elastic filament, above a critical strain they deviate from this simple elastic model, showing a softening response with increasingdeformations. A three-dimensional thin-shell model, in which the increased mechanical compli"},"claims":{"count":0,"items":[],"snapshot_sha256":"258153158e38e3291e3d48162225fcdb2d5a3ed65a07baac614ab91432fd4f57"},"source":{"id":"1807.08719","kind":"arxiv","version":1},"verdict":{"id":null,"model_set":{},"created_at":null,"strongest_claim":"","one_line_summary":"","pipeline_version":null,"weakest_assumption":"","pith_extraction_headline":""},"references":{"count":0,"sample":[],"resolved_work":0,"snapshot_sha256":"258153158e38e3291e3d48162225fcdb2d5a3ed65a07baac614ab91432fd4f57","internal_anchors":0},"formal_canon":{"evidence_count":0,"snapshot_sha256":"258153158e38e3291e3d48162225fcdb2d5a3ed65a07baac614ab91432fd4f57"},"author_claims":{"count":0,"strong_count":0,"snapshot_sha256":"258153158e38e3291e3d48162225fcdb2d5a3ed65a07baac614ab91432fd4f57"},"builder_version":"pith-number-builder-2026-05-17-v1"},"verdict_id":null},"signer":{"signer_id":"pith.science","signer_type":"pith_registry","key_id":"pith-v1-2026-05","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54"},"created_at":"2026-05-18T00:10:04Z","supersedes":[],"prev_event":null,"signature":{"signature_status":"signed_v1","algorithm":"ed25519","key_id":"pith-v1-2026-05","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54","signature_b64":"jqwgMgawTn2Iz32HPYOTdNf1KiLTtB6Evyor36vWQW9ZyKw4mBXbT4kKfNzf1z//sHMCoBvIPEKtHoEbcHkRAg==","signed_message":"open_graph_event_sha256_bytes","signed_at":"2026-06-02T14:16:58.869130Z"},"content_sha256":"b5edcf4b5f40d580cc4f2c0e61ac7577a6a71531d915e3598f296cf9e07ea2e3","schema_version":"1.0","event_id":"sha256:b5edcf4b5f40d580cc4f2c0e61ac7577a6a71531d915e3598f296cf9e07ea2e3"}],"timestamp_proofs":[],"mirror_hints":[{"mirror_type":"https","name":"Pith Resolver","base_url":"https://pith.science","bundle_url":"https://pith.science/pith/TOZC2JYUECSBABB4FSNA7P3MTQ/bundle.json","state_url":"https://pith.science/pith/TOZC2JYUECSBABB4FSNA7P3MTQ/state.json","well_known_bundle_url":"https://pith.science/.well-known/pith/TOZC2JYUECSBABB4FSNA7P3MTQ/bundle.json","status":"primary"}],"public_keys":[{"key_id":"pith-v1-2026-05","algorithm":"ed25519","format":"raw","public_key_b64":"stVStoiQhXFxp4s2pdzPNoqVNBMojDU/fJ2db5S3CbM=","public_key_hex":"b2d552b68890857171a78b36a5dccf368a953413288c353f7c9d9d6f94b709b3","fingerprint_sha256_b32_first128bits":"RVFV5Z2OI2J3ZUO7ERDEBCYNKS","fingerprint_sha256_hex":"8d4b5ee74e4693bcd1df2446408b0d54","rotates_at":null,"url":"https://pith.science/pith-signing-key.json","notes":"Pith uses this Ed25519 key to sign canonical record SHA-256 digests. Verify with: ed25519_verify(public_key, message=canonical_sha256_bytes, signature=base64decode(signature_b64))."}],"merge_version":"pith-open-graph-merge-v1","built_at":"2026-06-02T14:16:58Z","links":{"resolver":"https://pith.science/pith/TOZC2JYUECSBABB4FSNA7P3MTQ","bundle":"https://pith.science/pith/TOZC2JYUECSBABB4FSNA7P3MTQ/bundle.json","state":"https://pith.science/pith/TOZC2JYUECSBABB4FSNA7P3MTQ/state.json","well_known_bundle":"https://pith.science/.well-known/pith/TOZC2JYUECSBABB4FSNA7P3MTQ/bundle.json"},"state":{"state_type":"pith_open_graph_state","state_version":"1.0","pith_number":"pith:2018:TOZC2JYUECSBABB4FSNA7P3MTQ","merge_version":"pith-open-graph-merge-v1","event_count":2,"valid_event_count":2,"invalid_event_count":0,"equivocation_count":0,"current":{"canonical_record":{"metadata":{"abstract_canon_sha256":"3d4e56f7ddeb18472bc7593302363b75564b4d84f53a79440e6a8287b104d165","cross_cats_sorted":["q-bio.BM","q-bio.QM","q-bio.SC"],"license":"http://arxiv.org/licenses/nonexclusive-distrib/1.0/","primary_cat":"physics.bio-ph","submitted_at":"2018-07-23T17:00:57Z","title_canon_sha256":"1f520c67e85b1be981e4050352efc001a39271e1db33e603b88daed4b217dc79"},"schema_version":"1.0","source":{"id":"1807.08719","kind":"arxiv","version":1}},"source_aliases":[{"alias_kind":"arxiv","alias_value":"1807.08719","created_at":"2026-05-18T00:10:04Z"},{"alias_kind":"arxiv_version","alias_value":"1807.08719v1","created_at":"2026-05-18T00:10:04Z"},{"alias_kind":"doi","alias_value":"10.48550/arxiv.1807.08719","created_at":"2026-05-18T00:10:04Z"},{"alias_kind":"pith_short_12","alias_value":"TOZC2JYUECSB","created_at":"2026-05-18T12:32:53Z"},{"alias_kind":"pith_short_16","alias_value":"TOZC2JYUECSBABB4","created_at":"2026-05-18T12:32:53Z"},{"alias_kind":"pith_short_8","alias_value":"TOZC2JYU","created_at":"2026-05-18T12:32:53Z"}],"graph_snapshots":[{"event_id":"sha256:b5edcf4b5f40d580cc4f2c0e61ac7577a6a71531d915e3598f296cf9e07ea2e3","target":"graph","created_at":"2026-05-18T00:10:04Z","signer":{"key_id":"pith-v1-2026-05","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54","signer_id":"pith.science","signer_type":"pith_registry"},"payload":{"graph_snapshot":{"author_claims":{"count":0,"snapshot_sha256":"258153158e38e3291e3d48162225fcdb2d5a3ed65a07baac614ab91432fd4f57","strong_count":0},"builder_version":"pith-number-builder-2026-05-17-v1","claims":{"count":0,"items":[],"snapshot_sha256":"258153158e38e3291e3d48162225fcdb2d5a3ed65a07baac614ab91432fd4f57"},"formal_canon":{"evidence_count":0,"snapshot_sha256":"258153158e38e3291e3d48162225fcdb2d5a3ed65a07baac614ab91432fd4f57"},"paper":{"abstract_excerpt":"We use optical trapping to continuously bend an isolated microtubule while simultaneously measuring the applied force and the resulting filament strain, thus allowing us to determine its elastic properties over a wide range of applied strains. We find that, while in the low-strain regime, microtubules may be quantitatively described in terms of the classical Euler-Bernoulli elastic filament, above a critical strain they deviate from this simple elastic model, showing a softening response with increasingdeformations. A three-dimensional thin-shell model, in which the increased mechanical compli","authors_text":"Edvin Memet, Feodor Hilitski, L Mahadevan, Margaret A Morris, Walter J Schwenger, Zvonimir Dogic","cross_cats":["q-bio.BM","q-bio.QM","q-bio.SC"],"headline":"","license":"http://arxiv.org/licenses/nonexclusive-distrib/1.0/","primary_cat":"physics.bio-ph","submitted_at":"2018-07-23T17:00:57Z","title":"Microtubules soften due to cross-sectional flattening"},"references":{"count":0,"internal_anchors":0,"resolved_work":0,"sample":[],"snapshot_sha256":"258153158e38e3291e3d48162225fcdb2d5a3ed65a07baac614ab91432fd4f57"},"source":{"id":"1807.08719","kind":"arxiv","version":1},"verdict":{"created_at":null,"id":null,"model_set":{},"one_line_summary":"","pipeline_version":null,"pith_extraction_headline":"","strongest_claim":"","weakest_assumption":""}},"verdict_id":null}}],"author_attestations":[],"timestamp_anchors":[],"storage_attestations":[],"citation_signatures":[],"replication_records":[],"corrections":[],"mirror_hints":[],"record_created":{"event_id":"sha256:6ea04fa9a163482f4226b25790c548637e2bdcfed23c6c25d7a74476cf9ca8d3","target":"record","created_at":"2026-05-18T00:10:04Z","signer":{"key_id":"pith-v1-2026-05","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54","signer_id":"pith.science","signer_type":"pith_registry"},"payload":{"attestation_state":"computed","canonical_record":{"metadata":{"abstract_canon_sha256":"3d4e56f7ddeb18472bc7593302363b75564b4d84f53a79440e6a8287b104d165","cross_cats_sorted":["q-bio.BM","q-bio.QM","q-bio.SC"],"license":"http://arxiv.org/licenses/nonexclusive-distrib/1.0/","primary_cat":"physics.bio-ph","submitted_at":"2018-07-23T17:00:57Z","title_canon_sha256":"1f520c67e85b1be981e4050352efc001a39271e1db33e603b88daed4b217dc79"},"schema_version":"1.0","source":{"id":"1807.08719","kind":"arxiv","version":1}},"canonical_sha256":"9bb22d271420a410043c2c9a0fbf6c9c0f28d21aee70b41c2ea6d95c2ea9e2ae","receipt":{"algorithm":"ed25519","builder_version":"pith-number-builder-2026-05-17-v1","canonical_sha256":"9bb22d271420a410043c2c9a0fbf6c9c0f28d21aee70b41c2ea6d95c2ea9e2ae","first_computed_at":"2026-05-18T00:10:04.897537Z","key_id":"pith-v1-2026-05","kind":"pith_receipt","last_reissued_at":"2026-05-18T00:10:04.897537Z","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54","receipt_version":"0.3","signature_b64":"wyQKi+UgxRgE5WdPg0P5q5nK8ygo1P9y1O6CbrNy/AO6alXBlsDYqA7qR7pFMXb1Rl7VbUd/ZDQgKq7qr0cMDg==","signature_status":"signed_v1","signed_at":"2026-05-18T00:10:04.898213Z","signed_message":"canonical_sha256_bytes"},"source_id":"1807.08719","source_kind":"arxiv","source_version":1}}},"equivocations":[],"invalid_events":[],"applied_event_ids":["sha256:6ea04fa9a163482f4226b25790c548637e2bdcfed23c6c25d7a74476cf9ca8d3","sha256:b5edcf4b5f40d580cc4f2c0e61ac7577a6a71531d915e3598f296cf9e07ea2e3"],"state_sha256":"78f5891d3e0e331ef8eadd751abc122535e6f3b4090e3f916c73f593a428bdb1"},"bundle_signature":{"signature_status":"signed_v1","algorithm":"ed25519","key_id":"pith-v1-2026-05","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54","signature_b64":"LAmKnzOs/P+0WhN+MJaOkAWi5dOZv+V6SDi2trYicNrqKMDRtDCOR/BkAx8AJrNepIkRy8r+kHTsOVrpy6hPBQ==","signed_message":"bundle_sha256_bytes","signed_at":"2026-06-02T14:16:58.871177Z","bundle_sha256":"c0608e0a978bf146dab08e83a42c1145b54aadbce56fd743160e2b0bb930ec3d"}}