{"state_type":"pith_open_graph_state","state_version":"1.0","pith_number":"pith:2016:US5WQCX7LD7LBA7DWNR54ZBKRH","merge_version":"pith-open-graph-merge-v1","event_count":2,"valid_event_count":2,"invalid_event_count":0,"equivocation_count":0,"current":{"canonical_record":{"metadata":{"abstract_canon_sha256":"72c0abba6eefb06ba14aa59aec5c4fcb607a14d22278b64475cb913934410b7d","cross_cats_sorted":["physics.bio-ph"],"license":"http://arxiv.org/licenses/nonexclusive-distrib/1.0/","primary_cat":"q-bio.CB","submitted_at":"2016-06-02T20:38:14Z","title_canon_sha256":"a7a9dd75c2505e09d31e8dfcba87f929ff7ae1621cf0aa67e14001c3818d32dd"},"schema_version":"1.0","source":{"id":"1606.00863","kind":"arxiv","version":2}},"source_aliases":[{"alias_kind":"arxiv","alias_value":"1606.00863","created_at":"2026-05-18T00:54:20Z"},{"alias_kind":"arxiv_version","alias_value":"1606.00863v2","created_at":"2026-05-18T00:54:20Z"},{"alias_kind":"doi","alias_value":"10.48550/arxiv.1606.00863","created_at":"2026-05-18T00:54:20Z"},{"alias_kind":"pith_short_12","alias_value":"US5WQCX7LD7L","created_at":"2026-05-18T12:30:46Z"},{"alias_kind":"pith_short_16","alias_value":"US5WQCX7LD7LBA7D","created_at":"2026-05-18T12:30:46Z"},{"alias_kind":"pith_short_8","alias_value":"US5WQCX7","created_at":"2026-05-18T12:30:46Z"}],"graph_snapshots":[{"event_id":"sha256:a157e4e9c4a47d66da344e25c40d911d6d892a7566e7bc15e4f060e196c8d684","target":"graph","created_at":"2026-05-18T00:54:20Z","signer":{"key_id":"pith-v1-2026-05","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54","signer_id":"pith.science","signer_type":"pith_registry"},"payload":{"graph_snapshot":{"author_claims":{"count":0,"snapshot_sha256":"258153158e38e3291e3d48162225fcdb2d5a3ed65a07baac614ab91432fd4f57","strong_count":0},"builder_version":"pith-number-builder-2026-05-17-v1","claims":{"count":0,"items":[],"snapshot_sha256":"258153158e38e3291e3d48162225fcdb2d5a3ed65a07baac614ab91432fd4f57"},"formal_canon":{"evidence_count":0,"snapshot_sha256":"258153158e38e3291e3d48162225fcdb2d5a3ed65a07baac614ab91432fd4f57"},"paper":{"abstract_excerpt":"Cilia and flagella exhibit regular bending waves that perform mechanical work on the surrounding fluid, to propel cellular swimmers and pump fluids inside organisms. Here, we quantify a force-velocity relationship of the beating flagellum, by exposing flagellated \\emph{Chlamydomonas} cells to controlled microfluidic flows. A simple theory of flagellar limit-cycle oscillations, calibrated by measurements in the absence of flow, reproduces this relationship quantitatively. We derive a link between the chemo-mechanical efficiency of the flagellar beat and its ability to synchronize to oscillatory","authors_text":"Benjamin M. Friedrich, Christian Ruloff, Christian Wanger, Gary S. Klindt","cross_cats":["physics.bio-ph"],"headline":"","license":"http://arxiv.org/licenses/nonexclusive-distrib/1.0/","primary_cat":"q-bio.CB","submitted_at":"2016-06-02T20:38:14Z","title":"The load-response of the flagellar beat"},"references":{"count":0,"internal_anchors":0,"resolved_work":0,"sample":[],"snapshot_sha256":"258153158e38e3291e3d48162225fcdb2d5a3ed65a07baac614ab91432fd4f57"},"source":{"id":"1606.00863","kind":"arxiv","version":2},"verdict":{"created_at":null,"id":null,"model_set":{},"one_line_summary":"","pipeline_version":null,"pith_extraction_headline":"","strongest_claim":"","weakest_assumption":""}},"verdict_id":null}}],"author_attestations":[],"timestamp_anchors":[],"storage_attestations":[],"citation_signatures":[],"replication_records":[],"corrections":[],"mirror_hints":[],"record_created":{"event_id":"sha256:d03cfd41b303ed9b26b198951e0ce2daf1e89e9dc954c3fe9bda31c76074b4de","target":"record","created_at":"2026-05-18T00:54:20Z","signer":{"key_id":"pith-v1-2026-05","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54","signer_id":"pith.science","signer_type":"pith_registry"},"payload":{"attestation_state":"computed","canonical_record":{"metadata":{"abstract_canon_sha256":"72c0abba6eefb06ba14aa59aec5c4fcb607a14d22278b64475cb913934410b7d","cross_cats_sorted":["physics.bio-ph"],"license":"http://arxiv.org/licenses/nonexclusive-distrib/1.0/","primary_cat":"q-bio.CB","submitted_at":"2016-06-02T20:38:14Z","title_canon_sha256":"a7a9dd75c2505e09d31e8dfcba87f929ff7ae1621cf0aa67e14001c3818d32dd"},"schema_version":"1.0","source":{"id":"1606.00863","kind":"arxiv","version":2}},"canonical_sha256":"a4bb680aff58feb083e3b363de642a89c7a8ed2b754041838f202a7e96ef1b46","receipt":{"algorithm":"ed25519","builder_version":"pith-number-builder-2026-05-17-v1","canonical_sha256":"a4bb680aff58feb083e3b363de642a89c7a8ed2b754041838f202a7e96ef1b46","first_computed_at":"2026-05-18T00:54:20.327524Z","key_id":"pith-v1-2026-05","kind":"pith_receipt","last_reissued_at":"2026-05-18T00:54:20.327524Z","public_key_fingerprint":"8d4b5ee74e4693bcd1df2446408b0d54","receipt_version":"0.3","signature_b64":"EKQZwbr7PXDQQe4FnX9+IAibKusV6TdCJvsOxWDjrlrU4pzNC/EcQF2ZAR9NP9to+isaMVFge1j+Yj7ZAXlyDA==","signature_status":"signed_v1","signed_at":"2026-05-18T00:54:20.327970Z","signed_message":"canonical_sha256_bytes"},"source_id":"1606.00863","source_kind":"arxiv","source_version":2}}},"equivocations":[],"invalid_events":[],"applied_event_ids":["sha256:d03cfd41b303ed9b26b198951e0ce2daf1e89e9dc954c3fe9bda31c76074b4de","sha256:a157e4e9c4a47d66da344e25c40d911d6d892a7566e7bc15e4f060e196c8d684"],"state_sha256":"fe098707a555bc70864dec4a4fc37255b7584116e02062e35cf34491283e3c26"}