Spatially extended heterogeneous metacommunities can sustain long-lived chaotic fluctuations, and these fluctuations allow more species to coexist than the same system could at equilibrium.
Population variability in space and time.Trends in Ecology & Evolution , 15(11):460–464, November 2000
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Can endogenous fluctuations persist in high-diversity ecosystems?
Spatially extended heterogeneous metacommunities can sustain long-lived chaotic fluctuations, and these fluctuations allow more species to coexist than the same system could at equilibrium.