Chemotactic gradients above a threshold make multicellular clusters elongate perpendicular to the gradient and fragment, and this instability regulates cluster size and optimizes forward migration.
In this scenario, the probabilityp i for any particle to be in a particular layer is equal for all layers
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Chemotaxis-Driven Instabilities Govern Size, Shape and Migration Efficiency of Multicellular Clusters
Chemotactic gradients above a threshold make multicellular clusters elongate perpendicular to the gradient and fragment, and this instability regulates cluster size and optimizes forward migration.