For a ciliated planktonic organism modeled as a sphere in Stokes flow, surface shear measurements reveal the full strain rate, but horizontal vorticity is measurable only when the organism is bottom-heavy and tilts rapidly.
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Hydrodynamics of flow sensing in plankton
For a ciliated planktonic organism modeled as a sphere in Stokes flow, surface shear measurements reveal the full strain rate, but horizontal vorticity is measurable only when the organism is bottom-heavy and tilts rapidly.