High rank linear syzygies on low rank quadrics

We study the linear syzygies of a homogeneous ideal I in a polynomial ring S, focusing on the graded betti numbers b_(i,i+1) = dim_k Tor_i(S/I, k)_(i+1). For a variety X and divisor D with S = Sym(H^0(D)*), what conditions on D ensure that b_(i,i+1) is nonzero? Eisenbud has shown that a decomposition D = A + B such that A and B have at least two sections gives rise to determinantal equations (and corresponding syzygies) in I_X; and conjectured that if I_2 is generated by quadrics of rank at most 4, then the last nonvanishing b_(i,i+1) is a consequence of such equations. We describe obstructions to this conjecture and prove a variant. The obstructions arise from toric specializations of the Rees algebra of Koszul cycles, and we give an explicit construction of toric varieties with minimal linear syzygies of arbitrarily high rank. This gives one answer to a question posed by Eisenbud and Koh about specializations of syzygies.