Generation of second maximal subgroups and the existence of special primes

Let $G$ be a finite almost simple group. It is well known that $G$ can be generated by 3 elements, and in previous work we showed that 6 generators suffice for all maximal subgroups of $G$. In this paper we consider subgroups at the next level of the subgroup lattice - the so-called second maximal subgroups. We prove that with the possible exception of some families of rank 1 groups of Lie type, the number of generators of every second maximal subgroup of $G$ is bounded by an absolute constant. We also show that such a bound holds without any exceptions if and only if there are only finitely many primes $r$ for which there is a prime power $q$ such that $(q^r-1)/(q-1)$ is prime. The latter statement is a formidable open problem in Number Theory. Applications to random generation and polynomial growth are also given.