The formation and evolution of supermassive black holes and their host galaxies

We discuss constraints on the assembly history of supermassive black holes from the observed remnant black holes in nearby galaxies and from the emission caused by accretion onto these black holes. We also summarize the results of a specific model for the evolution of galaxies and their central black holes which traces their hierachical build-up in CDM-like cosmogonies. The model assumes (i) that black holes, ellipticals and starburts form during major mergers of galaxies (ii) that the gas fraction in galaxies decreases with decreasing redshift (iii) that the optical bright phase of a QSO lasts for about 10^7 years. The model succesfully reproduces the evolution of cold gas as traced by damped damped Lyman alpha systems, the evolution of optically bright QSOs, the remnant black hole mass distribution and the host galaxy luminosities of QSOs.