Lucas-type congruences for cyclotomic psi-coefficients

Let p be any prime and a be a positive integer. For nonnegative integers l,n and an integer r, the normalized cyclotomic $\psi$-coefficient $${n,r}_{l,p^a}:=p^{-[(n-p^{a-1}-lp^a)/(p^{a-1}(p-1))]} \sum_{k=r(mod p^a)}(-1)^k{n \choose k}{{(k-r)/p^a} \choose l}$$ is known to be an integer. In this paper, we show that this coefficient behaves like binomial coefficients and satisfies some Lucas-type congruences. This implies that a congruence of Wan is often optimal, and two conjectures of Sun and Davis are true.