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Paper Citation Record · LEDGER

BERTology Meets Biology: Interpreting Attention in Protein Language Models

As of 8 August 2026, this Paper Citation Record lists 0 of 0 outbound references and 8 inbound Pith citation observations for arXiv:2006.15222.

A citation records a reference. It does not transfer a finding from one paper to another.

pith.paper-citation-record.v1
2006.15222 v3

Coverage vector

measured 0 of 0 reference resolution

Typed states for the displayed outbound observations.

Source: paper_references, paper_reference_links

measured 8 of 8 standing notices

One-hop event checks from named stored sources.

Source: scholarly_work_events, retraction_status_cache, observed 2026-08-08T06:32:00.761636+00:00

measured 8 of 8 inbound itemization

Pith citing papers itemized under the disclosed page cap.

Source: paper_references, paper_reference_links, observed 2026-08-07T22:36:43.483055Z

measured 1 of 1 external citation measurements

A source-named dated measurement, never combined with another source.

Source: arxiv_reference, observed 2026-08-05T02:28:24.338817Z

Reference resolution

0 of 0 outbound references displayed

  • verified exact0
  • verified fuzzy0
  • unresolved0
  • parse uncertain0
  • malformed identifier0
  • metadata mismatch0

External citation measurements

28
arxiv_reference, observed 2026-08-05T02:28:24.338817Z

Outbound references

No outbound reference observations are available for this paper version.

Pith citing papers

Observation 04eb73f2-e466-4808-8a63-8d471cd578fe · inbound

Rethinking Attention with Performers cites this paper.

Rethinking Attention with Performers BERTology Meets Biology: Interpreting Attention in Protein Language Models

Reference 157

Resolution
metadata mismatch
arxiv_id, observed 2026-05-12T09:16:14.519510Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-08T06:32:00.761636+00:00.

source=arxiv_source observed=2026-05-12T09:16:14.336570Z digest=sha256:94880f7208d4c561574d34c2a34be7888d25f5a20e5d4ab76024c280f9d7e108

Observation 9f9aee10-4a8a-4eb8-b176-ad2117c9059a · inbound

Recent Advances in Generative AI for Healthcare Applications cites this paper.

Recent Advances in Generative AI for Healthcare Applications BERTology Meets Biology: Interpreting Attention in Protein Language Models

Reference 27

Resolution
verified exact
arxiv_id, observed 2026-05-24T06:29:00.462060Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-08T06:32:00.761636+00:00.

source=pdf_text observed=2026-05-24T06:26:31.708511Z digest=sha256:ad1d5d806b7fe36bcf6a9828f113afcfe9e1cb9a8999f66590c63fb0f0fa6642

Observation 642a30c2-00b0-41fa-9fce-f5b6c9885a21 · inbound

Interpreting and Steering Protein Language Models through Sparse Autoencoders cites this paper.

Interpreting and Steering Protein Language Models through Sparse Autoencoders BERTology Meets Biology: Interpreting Attention in Protein Language Models

Reference 2017

Resolution
unresolved
no resolver link, observed 2026-08-07T22:36:43.483055Z

Source-reported events for the cited work

Unavailable: canonical work link unavailable.

source=pdf_text observed=2026-08-07T22:36:43.483055Z digest=sha256:b982169df138770edda006e03fcdb9c36708fe1e5a2229e7850fc00d78664203

Observation 4e927c67-1829-4455-9fc0-2c20b053f620 · inbound

Enhancing Safe and Controllable Protein Generation via Knowledge Preference Optimization cites this paper.

Enhancing Safe and Controllable Protein Generation via Knowledge Preference Optimization BERTology Meets Biology: Interpreting Attention in Protein Language Models

Reference 32

Resolution
unresolved
no resolver link, observed 2026-08-06T17:26:45.473419Z

Source-reported events for the cited work

Unavailable: canonical work link unavailable.

source=arxiv_source observed=2026-08-06T17:26:45.473419Z digest=sha256:76600f0caa1f93c1882c4d74b3e3a8e8669461822405c1c97df6721f950e59a0

Observation 40f59fe9-8ee2-4b18-a167-c6a166372103 · inbound

Directed Evolution of Proteins via Bayesian Optimization in Embedding Space cites this paper.

Directed Evolution of Proteins via Bayesian Optimization in Embedding Space BERTology Meets Biology: Interpreting Attention in Protein Language Models

Reference 23

Resolution
unresolved
no resolver link, observed 2026-08-05T05:45:30.038663Z

Source-reported events for the cited work

Unavailable: canonical work link unavailable.

source=pdf_text observed=2026-08-05T05:45:30.038663Z digest=sha256:c4c6f325f305e0ebbd8094170c38b1a9c2a147a7a14b2217ce3cf2fed7150780

Observation 1f8c7576-16c7-4db0-bdfb-317512105120 · inbound

Retrieval and competition: how a protein foundation model starts a protein cites this paper.

Retrieval and competition: how a protein foundation model starts a protein BERTology Meets Biology: Interpreting Attention in Protein Language Models

Reference 7

Resolution
metadata mismatch
arxiv_id, observed 2026-05-21T00:13:53.256660Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-08T06:32:00.761636+00:00.

source=pdf_text observed=2026-05-21T00:09:22.402365Z digest=sha256:30a1c364732145278a86b43624806796c1cbf6d57ec8f6ce213885cfd646de1b

Observation eb3de4c0-5276-4723-a1ef-7979c23a6a20 · inbound

Retrieval and competition: how a protein foundation model starts a protein cites this paper.

Retrieval and competition: how a protein foundation model starts a protein BERTology Meets Biology: Interpreting Attention in Protein Language Models

Reference 7

Resolution
metadata mismatch
arxiv_id, observed 2026-07-01T00:25:10.454039Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-08T06:32:00.761636+00:00.

source=pdf_text observed=2026-06-30T23:58:11.169748Z digest=sha256:24f27eb1f0ddd71d583ceea04636162bef03daae5e5aeba0699ba403c21c4e0c

Observation 8ba3225c-1616-4875-baf9-0b3dfa1209cb · inbound

Evolution-Aware MSA Reasoning for Subsampling via Factor Graphs cites this paper.

Evolution-Aware MSA Reasoning for Subsampling via Factor Graphs BERTology Meets Biology: Interpreting Attention in Protein Language Models

Reference 46

Resolution
unresolved
no resolver link, observed 2026-08-01T05:12:53.254465Z

Source-reported events for the cited work

Unavailable: canonical work link unavailable.

source=pdf_text observed=2026-08-01T05:12:53.254465Z digest=sha256:7114d2bc202144f0bc1a02e361aa293b3af6a5fa7fcc81659c81491eb999eb48