Spanning trees in graphs without large bipartite holes

We show that for any $\varepsilon>0$ and $\Delta\in\mathbb{N}$, there exists $\alpha>0$ such that for sufficiently large $n$, every $n$-vertex graph $G$ satisfying that $\delta(G)\geq\varepsilon n$ and $e(X, Y)>0$ for every pair of disjoint vertex sets $X, Y\subseteq V(G)$ of size $\alpha n$ contains all spanning trees with maximum degree at most $\Delta$. This strengthens a result of B\"ottcher et al.