Typed states for the displayed outbound observations.
Source: paper_references, paper_reference_links, observed 2026-08-15T20:53:10.696679Z
Paper Citation Record · LEDGER
As of 17 August 2026, this Paper Citation Record lists 62 of 62 outbound references and 1 inbound Pith citation observation for arXiv:2505.11812.
A citation records a reference. It does not transfer a finding from one paper to another.
Typed states for the displayed outbound observations.
Source: paper_references, paper_reference_links, observed 2026-08-15T20:53:10.696679Z
One-hop event checks from named stored sources.
Source: scholarly_work_events, retraction_status_cache, observed 2026-08-17T06:30:58.91139+00:00
Pith citing papers itemized under the disclosed page cap.
Source: paper_references, paper_reference_links, observed 2026-05-18T08:27:36.890001Z
A source-named dated measurement, never combined with another source.
Source: arxiv_reference, observed 2026-05-18T08:31:06.984753Z
62 of 62 outbound references displayed
External citation measurements
No source-named external measurement is stored.
Observation d42a0525-f7ee-4237-8bcc-bbeb08ab5264 · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins Highly accurate protein structure prediction with AlphaFold.Nature, 596(7873):583–589, 2021
Reference 1
Source-reported events for the cited work
Unavailable: canonical work link unavailable.
Observation d7c4754c-2d06-43ae-b041-ed884908e35e · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins Accurate structure prediction of biomolecular interactions with AlphaFold 3.Nature, pages 1–3, 2024
Reference 2
Source-reported events for the cited work
Unavailable: canonical work link unavailable.
Observation 79a67369-7b4b-40d7-8227-296897f77826 · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins Machine learning- aided engineering of hydrolases for PET depolymerization.Nature, 604(7907):662–667, 2022
Reference 3
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.
Observation e93f819d-af2d-46d7-a735-6ef8e05daf20 · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins A conditional protein diffusion model generates artificial programmable endonuclease sequences with enhanced activity.Cell Discovery, 10(1):95, 2024
Reference 4
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.
Observation 07ea00db-fac6-4497-b8b2-2818bef46822 · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins Retrieval-Enhanced Mutation Mastery: Augmenting Zero-Shot Prediction of Protein Language Model
Reference 5
Source-reported events for the cited work
Unavailable: canonical work link unavailable.
Observation 9e4eea15-dbe9-4721-a7cc-f157940818c8 · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins Enzyme function prediction using contrastive learning.Science, 379(6639):1358–1363, 2023
Reference 6
Source-reported events for the cited work
Unavailable: canonical work link unavailable.
Observation 134d9d99-c28e-45a3-90f0-c13dcbad400a · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins Enhancing efficiency of protein language models with minimal wet-lab data through few-shot learning.Nature Communications, 15(1):5566, 2024
Reference 7
Source-reported events for the cited work
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Observation 06328444-065b-4b59-b2c7-19f299b6a66f · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins CATH – a hierarchic classification of protein domain structures.Structure, 5(8):1093–1109, 1997
Reference 8
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.
Observation 4e1c78b3-0058-4308-ad89-550ed48c53a1 · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins Alphafold protein structure database: massively expanding the structural coverage of protein-sequence space with high-accuracy models.Nucleic Acids Research, 50(D1):D439–D444, 2022
Reference 9
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.
Observation 0d143272-4cac-4ff3-a9da-713f35414f2e · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins UniProt: the universal protein knowledgebase in 2025.Nucleic Acids Research, 53(D1):D609–D617, 2025
Reference 10
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.
Observation ec803221-e323-4541-bbbc-f0cac29cd481 · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins Simple, efficient, and scalable structure-aware adapter boosts protein language models.Journal of Chemical Information and Modeling, 2024
Reference 11
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.
Observation bbe1b82c-0375-40cb-a11b-21b612466096 · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins Immunogenicity prediction with dual attention enables vaccine target selection
Reference 12
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.
Observation 4803cbba-ba25-47ac-8db1-cfda43528f38 · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins Large-scale prediction of human protein- protein interactions from amino acid sequence based on latent topic features.Journal of Proteome Research, 9(10):4992–5001, 2010
Reference 13
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.
Observation 7a477de0-be9c-4e1c-82d8-0f751b9be89e · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins Unresolved cited work
Reference 14
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.
Observation dc396b4b-9416-445e-ad3c-9d251a896523 · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins Skempi 2.0: an updated benchmark of changes in protein–protein binding energy, kinetics and thermodynamics upon mutation.Bioinformatics, 35(3):462–469, 2019
Reference 15
Source-reported events for the cited work
Unavailable: canonical work link unavailable.
Observation ac738482-91fe-4e4b-af35-7e1ea18d7288 · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins A comprehensive dataset of protein-protein interactions and ligand binding pockets for advancing drug discovery.Scientific Data, 11(1):402, 2024
Reference 16
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.
Observation e4c043f5-26b9-4f44-9900-1b2d0e06d9f1 · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins Quan- titative missense variant effect prediction using large-scale mutagenesis data.Cell Systems, 6(1):116–124, 2018
Reference 17
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.
Observation 1c86f260-bda3-4ae3-94fd-eae7e36fa1da · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins Deep generative models of genetic variation capture the effects of mutations.Nature Methods, 15(10):816–822, 2018
Reference 18
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.
Observation 158348fb-a0b4-488f-b9fe-938349fbdac2 · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins ProteinGym: large-scale benchmarks for protein fitness prediction and design
Reference 19
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.
Observation 44c97d87-99d3-41d8-9416-0157f1e6c50e · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins Venusmuthub: a systematic evaluation of protein mutation effect predictors on small-scale experimental data.Acta Pharmaceutica Sinica B, 2025
Reference 20
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.
Observation 71a81df9-1fa8-4246-b218-f30001b4181e · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins Discovering functionally important sites in proteins.Nature communications, 14(1):4175, 2023
Reference 21
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.
Observation 435a7edb-d27f-4040-bf9e-0687264c5046 · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins Predicting protein function from sequence and structure.Nature reviews molecular cell biology, 8(12):995–1005, 2007
Reference 22
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.
Observation 49e2d6ac-33c7-48fb-b960-3b863b71b97a · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins Deep- bce: evaluation of deep learning models for identification of immunogenic b-cell epitopes
Reference 23
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.
Observation 171b7c4f-f73e-4dda-ada8-84369f85df84 · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins InterPro in 2022.Nucleic Acids Research, 51(D1):D418–D427, 2023
Reference 24
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.
Observation 9b974618-98be-4c3b-a753-32aa003e359a · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins BioLiP: a semi-manually curated database for biologically relevant ligand–protein interactions.Nucleic Acids Research, 41(D1):D1096– D1103, 2012
Reference 25
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.
Observation b50dd037-356e-4e43-b3d1-08919d2f23bb · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins SAbDab: the structural antibody database.Nucleic Acids Research, 42(D1):D1140–D1146, 2014
Reference 26
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.
Observation ecc5e8da-18a5-4054-aebc-3bfc9a29f06e · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins Unresolved cited work
Reference 27
Source-reported events for the cited work
Unavailable: canonical work link unavailable.
Observation 22713993-2fb4-4c07-bfaa-2dc106a08437 · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins Prottrans: Toward understanding the language of life through self-supervised learning.IEEE Transactions on Pattern Analysis and Machine Intelligence, 44(10):7112–7127, 2021
Reference 28
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.
Observation f4d175f3-18ab-4915-908b-9c85bc157171 · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins Ankh: Optimized Protein Language Model Unlocks General-Purpose Modelling
Reference 29
Source-reported events for the cited work
Unavailable: canonical work link unavailable.
Observation f3e751bc-2982-4d07-9f07-77c005cdcc34 · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins ProstT5: Bilingual language model for protein sequence and structure.bioRxiv, pages 2023–07, 2023
Reference 30
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.
Observation 1c7d8132-111e-44f3-881f-865ad5db25a9 · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins Evolutionary-scale prediction of atomic-level protein structure with a language model.Science, 379(6637):1123–1130, 2023
Reference 31
Source-reported events for the cited work
Unavailable: canonical work link unavailable.
Observation 6c71a887-21ab-4e2a-b217-25f7f677c86f · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins Protein remote homology detection and structural alignment using deep learning
Reference 32
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.
Observation 85e0a84e-d608-4bfe-aa0b-e58d046f2f2a · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins SaProt: protein language modeling with structure-aware vocabulary
Reference 33
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.
Observation e038233d-ef38-4642-be5c-98f43a228cda · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins Semantical and geo- metrical protein encoding toward enhanced bioactivity and thermostability.eLife, 13:RP98033, may 2025
Reference 34
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.
Observation 89f15533-4e03-4df9-9258-5c2e1b79b301 · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins Masked inverse folding with sequence transfer for protein representation learning.Protein Engineering, Design and Selection, 36:gzad015, 2023
Reference 35
Source-reported events for the cited work
Unavailable: canonical work link unavailable.
Observation 53cdceb7-f84d-4949-9bd8-5d7e074373c5 · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins Learning inverse folding from millions of predicted structures
Reference 36
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.
Observation be00b58d-8103-46fb-b1a6-65451ca24d8b · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins Learning from protein structure with geometric vector perceptrons
Reference 37
Source-reported events for the cited work
Unavailable: canonical work link unavailable.
Observation ae1e4a57-c5d9-4986-b36f-2dfc1fa2a90b · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins Basic local alignment search tool.Journal of Molecular Biology, 215(3):403–410, 1990
Reference 38
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.
Observation 249429ea-6f68-4404-8ab0-e02b1f930108 · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins TM-align: a protein structure alignment algorithm based on the tm-score.Nucleic Acids Research, 33(7):2302–2309, 2005
Reference 39
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.
Observation 9b96d819-0184-4253-acd1-0abee1dc665a · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins Fast and accurate protein structure search with Foldseek.Nature Biotechnology, 42(2):243–246, 2024
Reference 40
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.
Observation 7d6fb224-19af-4ba6-bf51-7ad9c479c973 · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins VenusFactory: A Unified Platform for Protein Engineering Data Retrieval and Language Model Fine-Tuning
Reference 41
Source-reported events for the cited work
Unavailable: canonical work link unavailable.
Observation 93d21a52-5d66-41de-b79e-12f4edb493b7 · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins Unresolved cited work
Reference 42
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.
Observation acdd540a-508c-404b-846f-d4be399c0a1c · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins BioPython: freely available python tools for computational molecular biology and bioinformatics.Bioinformatics, 25(11):1422, 2009
Reference 43
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.
Observation 3b52f3e5-ab65-4cc5-9e55-070a445438ea · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins MMseqs2 enables sensitive protein sequence searching for the analysis of massive data sets.Nature Biotechnology, 35(11):1026–1028, 2017
Reference 44
Source-reported events for the cited work
Unavailable: canonical work link unavailable.
Observation 7fdb4247-6985-465d-a5f8-5cc9982636e0 · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins Unresolved cited work
Reference 45
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.
Observation 8e54af15-a0cc-4019-b834-29b1db0f055d · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins Decoupled Weight Decay Regularization
Reference 46
Source-reported events for the cited work
Unavailable: canonical work link unavailable.
Observation 6e5058f6-0ac5-4914-b400-744e8d4bb0cd · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins Evaluating protein transfer learning with TAPE.Advances in Neural Information Processing Systems, 32, 2019
Reference 47
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.
Observation 07977bc8-5694-4c21-859f-434f116c9eaa · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins ProteinNet: a standardized data set for machine learning of protein structure.BMC Bioinformatics, 20:1–10, 2019
Reference 48
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.
Observation 771a964f-bdd0-4df6-829a-a066c24fc9c3 · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins PEER: A comprehensive and multi-task benchmark for protein sequence understanding
Reference 49
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.
Observation 6f395a68-438e-4cd2-bcde-9626b986befe · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins PETA: evaluating the impact of protein transfer learning with sub-word tokenization on downstream applications.Journal of Cheminformatics, 16(1):92, 2024
Reference 50
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.
Observation f02469a6-f0d8-41d6-b85a-2899190fce20 · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins ProteinGLUE multi-task benchmark suite for self-supervised protein modeling
Reference 51
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.
Observation 41998c09-0ed2-4de5-87d7-7f4a8120ea07 · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins DeepLoc: prediction of protein subcellular localization using deep learning
Reference 52
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.
Observation d24de8f1-40c4-49ce-adc1-bc96eccb8d61 · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins DeepSol: a deep learning framework for sequence-based protein solubility prediction
Reference 53
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.
Observation 72401836-fdf9-4a7f-855a-3915b11442b5 · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins FLIP: Benchmark tasks in fitness landscape inference for proteins
Reference 54
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.
Observation 1d6ee837-c069-4f4b-8504-2dd113ddb297 · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins ProteinShake: Building datasets and benchmarks for deep learning on protein structures
Reference 55
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.
Observation 02f48977-f3f3-4c00-aa23-5fbc2fb991c7 · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins Using support vector machine combined with auto covariance to predict protein–protein interactions from protein sequences
Reference 56
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.
Observation 5fb65a95-cd70-4ee6-8c85-46f8f1518c42 · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins Deciphering interaction fingerprints from protein molecular surfaces using geometric deep learning.Nature Methods, 17(2):184–192, 2020
Reference 57
Source-reported events for the cited work
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Observation e638c03b-743d-49ed-af62-8f10d26dc6df · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins DIPS-plus: The enhanced database of interacting protein structures for interface prediction.Scientific Data, 10(1):509, 2023
Reference 58
Source-reported events for the cited work
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Observation 6de9e84c-7f6d-4d4a-87a4-56b882374b6e · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins A comprehensive dataset of protein-protein interactions and ligand binding pockets for advancing drug discovery.Scientific Data, 11(1):402, 2024
Reference 59
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.
Observation a42f59df-ff1e-4bef-9ec1-33d030fcbcfb · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins The PDBbind database: methodologies and updates.Journal of Medicinal Chemistry, 48(12):4111–4119, 2005
Reference 60
Source-reported events for the cited work
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Observation 09aaa3c4-c990-4a4d-b417-dcc080486072 · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins ATOM3D: Tasks On Molecules in Three Dimensions
Reference 61
Source-reported events for the cited work
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Observation f541a08b-7abd-4cdd-868c-51e4dd11b845 · outbound
VenusX: Unlocking Fine-Grained Functional Understanding of Proteins Task” indicates evaluation scope: “All
Reference 62
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.
Observation 41b94bc0-a7e4-4c2f-9a50-52921d4e12b4 · inbound
Fast and Interpretable Protein Substructure Alignment via Optimal Transport VenusX: Unlocking Fine-Grained Functional Understanding of Proteins
Reference 22
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.