Typed states for the displayed outbound observations.
Source: paper_references, paper_reference_links, observed 2026-08-07T11:36:32.385533Z
Paper Citation Record · LEDGER
As of 8 August 2026, this Paper Citation Record lists 59 of 59 outbound references and 0 inbound Pith citation observations for arXiv:2506.04264.
A citation records a reference. It does not transfer a finding from one paper to another.
Typed states for the displayed outbound observations.
Source: paper_references, paper_reference_links, observed 2026-08-07T11:36:32.385533Z
One-hop event checks from named stored sources.
Source: scholarly_work_events, retraction_status_cache, observed 2026-08-07T06:34:17.273281+00:00
Pith citing papers itemized under the disclosed page cap.
Source: paper_references, paper_reference_links
A source-named dated measurement, never combined with another source.
Source: cited_works
59 of 59 outbound references displayed
External citation measurements
No source-named external measurement is stored.
Observation 57895fb5-7525-452a-914c-1f6aa0f9d9bb · outbound
Direct reciprocity in asynchronous interactions Spatial dilemmas of diffusible public goods.Elife, 2:e01169, 2013
Reference 1
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation 1ecef70e-0bc3-4ef1-8f69-6a2097702d40 · outbound
Direct reciprocity in asynchronous interactions Spatial structure, cooperation and competition in biofilms.Nature Reviews Microbiology, 14:589–600, 2016
Reference 2
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation 751c536c-0d5d-42f6-88e8-a6995722d1b2 · outbound
Direct reciprocity in asynchronous interactions Reciprocal food sharing in the vampire bat.Nature, 308:181–184, 1984
Reference 3
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation 6a25c37e-6497-49a0-822c-ffb56cdc7142 · outbound
Direct reciprocity in asynchronous interactions Unresolved cited work
Reference 4
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation c5dcdc0f-c504-414f-bb50-3669a198797d · outbound
Direct reciprocity in asynchronous interactions Chimpanzees share food for many reasons: the role of kinship, reciprocity, social bonds and harassment on food transfers.Animal Behaviour, 85:941–947, 2013
Reference 5
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation 5c84c855-bd45-4ab9-a3c1-9d0e4167f327 · outbound
Direct reciprocity in asynchronous interactions How is human cooperation different?Philosophical Transactions of the Royal Society B, 365:2663–2674, 2010
Reference 6
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation af7d9473-cb1f-43e6-8915-69dcc1f13685 · outbound
Direct reciprocity in asynchronous interactions Human cooperation.Trends in Cognitive Sciences, 17:413–425, 2013
Reference 7
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation b4a37b75-da93-4ddb-bae3-dcf1e4fddcf9 · outbound
Direct reciprocity in asynchronous interactions The tragedy of the commons: the population problem has no technical solution; it requires a fundamental extension in morality.Science, 162:1243–1248, 1968
Reference 8
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation 167ed681-7280-4b46-92b4-8f33a841217f · outbound
Direct reciprocity in asynchronous interactions Super-additive coop- eration.Nature, 626:1034–1041, 2024
Reference 9
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation 95a26e93-4d2b-4636-a4ac-87e008405931 · outbound
Direct reciprocity in asynchronous interactions Cooper- ating with the future.Nature, 511:220–223, 2014
Reference 10
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation af6f757a-41b5-4db1-8c1e-458c04060236 · outbound
Direct reciprocity in asynchronous interactions The evolution of eusociality
Reference 11
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation 4d7422a7-0ae0-4e9c-83f7-83536de047d4 · outbound
Direct reciprocity in asynchronous interactions Altruism in viscous populations—an inclusive fitness model.Evolutionary Ecology, 6:352–356, 1992
Reference 12
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation 7719cef3-0e66-4710-9e61-5f7ffee6cfce · outbound
Direct reciprocity in asynchronous interactions A theoretical basis for measures of kin selection in subdivided populations: finite populations and localized dispersal.Journal of Evolutionary Biology, 13:814–825, 2000
Reference 13
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation de0be6f8-ac6f-4fca-9c1f-569bfebd3971 · outbound
Direct reciprocity in asynchronous interactions Evolu- tionary instability of selfish learning in repeated games.PNAS nexus, 1:pgac141, 2022
Reference 14
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation cec80ce1-c1b2-452f-b5b1-45ac1f699a99 · outbound
Direct reciprocity in asynchronous interactions Mutation enhances cooperation in direct reciprocity.Proceedings of the National Academy of Sciences, 120:e2221080120, 2023
Reference 15
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation c9109010-9c4b-4536-9f59-456a37b5318b · outbound
Direct reciprocity in asynchronous interactions Indirect reciprocity with optional interactions.Journal of Theoretical Biology, 365:1–11, 2015
Reference 16
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation 42a0afca-0a58-4958-b52e-0d78a1fa1bb2 · outbound
Direct reciprocity in asynchronous interactions A solution for private assessment in indirect reciprocity using solitary observation.Journal of Theoretical Biology, 455:7–15, 2018
Reference 17
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation e417cd05-01c5-4121-920f-d384ac9f3c60 · outbound
Direct reciprocity in asynchronous interactions The leading eight: social norms that can maintain coop- eration by indirect reciprocity.Journal of Theoretical Biology, 239:435–444, 2006
Reference 18
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation ea130fa6-047a-4a79-a9df-bb47e8590273 · outbound
Direct reciprocity in asynchronous interactions A simple rule for the evolution of cooperation on graphs and social networks.Nature, 441:502–505, 2006
Reference 19
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation fc1106b8-21db-4f05-91a9-90aae48e79b5 · outbound
Direct reciprocity in asynchronous interactions Evolutionary dynamics on any population struc- ture.Nature, 544:227–230, 2017
Reference 20
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation 91c60ca3-d3d6-4f64-83e7-0999940bd309 · outbound
Direct reciprocity in asynchronous interactions Strategy evolution on dynamic networks
Reference 21
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation cfdfc9c1-e9d2-4501-80e6-c31bae9601d1 · outbound
Direct reciprocity in asynchronous interactions Strategy evolution on higher- order networks.Nature Computational Science, 4:274–284, 2024
Reference 22
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation 7bb6f561-bbff-4588-96dd-97a56a2df765 · outbound
Direct reciprocity in asynchronous interactions Evolution of prosocial be- haviours in multilayer populations.Nature Human Behaviour, 6:338–348, 2022
Reference 23
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation 449518dd-ce25-4164-8cdd-472cbfd8f32d · outbound
Direct reciprocity in asynchronous interactions Evolution of cooperation with asymmetric social interactions.Proceedings of the National Academy of Sciences, 119:e2113468118, 2022
Reference 24
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation bbf0c64c-274c-4ddb-b30c-ec5c9cff6de3 · outbound
Direct reciprocity in asynchronous interactions Social goods dilemmas in hetero- geneous societies.Nature Human Behaviour, 4:819–831, 2020
Reference 25
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation 832fb68f-ef17-4b1c-a9eb-b4128e21ceb4 · outbound
Direct reciprocity in asynchronous interactions Evolution of cooperation on temporal networks.Nature communications, 11:2259, 2020
Reference 26
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation 12bf45cf-9623-491a-995e-403ad613c6c3 · outbound
Direct reciprocity in asynchronous interactions Evolutionary dynamics within and among competing groups.Proceedings of the National Academy of Sciences, 120:e2216186120, 2023
Reference 27
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation 202727e2-559d-4e62-b3e9-efd86608cdee · outbound
Direct reciprocity in asynchronous interactions Evolution of cooperation by generalized reciprocity.Proceedings of the Royal Society B, 272:1115–1120, 2005
Reference 28
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation 1360b789-84fc-4c54-b46f-5a0c6d014577 · outbound
Direct reciprocity in asynchronous interactions Emergence of cooperation and evolutionary stability in finite populations.Nature, 428:646–650, 2004
Reference 29
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation 021f8a6a-a1cd-47d3-8e54-ee4eaf86d0cd · outbound
Direct reciprocity in asynchronous interactions Memory-n strategies of direct reciprocity.Proceedings of the National Academy of Sci- ences, 114:4715–4720, 2017
Reference 30
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation 4cd21095-e05d-4a44-87ac-a2f8f7380e26 · outbound
Direct reciprocity in asynchronous interactions Compar- ing reactive and memory-one strategies of direct reciprocity.Scientific Reports, 6:25676, 2016
Reference 31
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation 6c3641e5-41b9-440b-b609-adb61fb6b0a4 · outbound
Direct reciprocity in asynchronous interactions Adaptive dynamics of memory- one strategies in the repeated donation game.PLoS Computational Biology, 19:e1010987, 2023
Reference 32
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation c805f163-130e-49bf-8bfb-f3c1c23b6858 · outbound
Direct reciprocity in asynchronous interactions Cooperation and con- trol in multiplayer social dilemmas.Proceedings of the National Academy of Sciences, 111:16425–16430, 2014
Reference 33
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation 4895cf8f-eefa-4f82-8588-fe7eb5e409f6 · outbound
Direct reciprocity in asynchronous interactions Outlearning extortioners: unbending strategies can foster reciprocal fairness and cooperation.PNAS nexus, 2:pgad176, 2023
Reference 34
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation b154434d-0a12-4d69-8b42-386a95004202 · outbound
Direct reciprocity in asynchronous interactions Partners and rivals in direct reciprocity.Nature Human Behaviour, 2:469–477, 2018
Reference 35
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation 57616a7d-3961-4402-9f86-5402c828c696 · outbound
Direct reciprocity in asynchronous interactions Effects of increasing the number of players and memory size in the iterated prisoner’s dilemma: a numerical approach.Proceedings of the Royal Society of London
Reference 36
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation 72cdc34e-bbba-46e1-ba77-4bb18c903763 · outbound
Direct reciprocity in asynchronous interactions Crosstalk in concurrent repeated games impedes direct reciprocity and requires stronger levels of forgiveness.Nature Communications, 9:555, 2018
Reference 37
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation cc7789e3-1208-4617-894a-8ae9769c1609 · outbound
Direct reciprocity in asynchronous interactions From extortion to generosity, evolution in the it- erated prisoner’s dilemma.Proceedings of the National Academy of Sciences, 110:15348– 15353, 2013
Reference 38
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation e3cd4786-4d6a-4e0d-9ce8-0d7234d071fa · outbound
Direct reciprocity in asynchronous interactions Evo- lution of all-or-none strategies in repeated public goods dilemmas.PLoS Computational Biology, 10:e1003945, 2014
Reference 39
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation 169c9c1a-5210-49d8-a041-806032ca61d7 · outbound
Direct reciprocity in asynchronous interactions The evolution of cooperation.Science, 211:1390–1396, 1981
Reference 40
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation 587d6156-e8a3-4e52-95fb-04b8a6b92c16 · outbound
Direct reciprocity in asynchronous interactions Tit for tat in heterogeneous populations.Nature, 355:250–253, 1992
Reference 41
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation 0d6b8604-ba54-4c15-9ce9-bca46e489256 · outbound
Direct reciprocity in asynchronous interactions A strategy of win-stay, lose-shift that outperforms tit-for-tat in the Prisoner’s Dilemma game.Nature, 364:56–58, 1993
Reference 42
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation 681b0451-a29c-447e-9392-2dbe33e9b964 · outbound
Direct reciprocity in asynchronous interactions Iterated prisoner’s dilemma contains strategies that dominate any evolutionary opponent.Proceedings of the National Academy of Sciences, 109:10409–10413, 2012
Reference 43
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation 5caa90ca-6d46-4858-9ed6-f79e30fd2b4c · outbound
Direct reciprocity in asynchronous interactions What you gotta know to play good in the iterated prisoner’s dilemma.Games, 6:175–190, 2015
Reference 44
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation 296416f0-b7c1-41ca-8e21-5e93411a1f78 · outbound
Direct reciprocity in asynchronous interactions Functional significance of social grooming in primates.Folia Prima- tologica, 57:121–131, 1991
Reference 45
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation 39bf3d37-1ada-4489-b74b-0a84039f5272 · outbound
Direct reciprocity in asynchronous interactions Is sentinel behaviour safe? An experimental investigation.Animal Behaviour, 85:137–142, 2013
Reference 46
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation cbf3d4d5-7b29-4513-89ef-ee6cbefc47f4 · outbound
Direct reciprocity in asynchronous interactions The alternating prisoner’s dilemma.Journal of Theoretical Biology, 168:219–226, 1994
Reference 47
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation 5b3e640c-9a94-4bd8-8e00-c2c59b9741e4 · outbound
Direct reciprocity in asynchronous interactions The prisoner’s dilemma without synchrony.Proceedings of the Royal Society of London
Reference 48
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation 92b85eee-1f2b-4a6c-85b6-bc716ac7ff3d · outbound
Direct reciprocity in asynchronous interactions Forgiver triumphs in alternating prisoner’s dilemma.PLoS One, 8:e80814, 2013
Reference 49
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation b1a4c85a-0453-4e79-a4dc-75e225a35d1d · outbound
Direct reciprocity in asynchronous interactions Autocratic strategies for alternating games.Theo- retical Population Biology, 113:13–22, 2017
Reference 50
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation 7bf7d7b8-779b-4310-861a-06a6840c6a57 · outbound
Direct reciprocity in asynchronous interactions Cooperation in alternating interac- tions with memory constraints.Nature Communications, 13:737, 2022
Reference 51
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation 13497580-d86d-444d-b1c4-8220a9c56727 · outbound
Direct reciprocity in asynchronous interactions Evolutionary games with environ- mental feedbacks.Nature Communications, 11:915, 2020
Reference 52
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation 8690a285-ab07-42c0-9550-b61d48092761 · outbound
Direct reciprocity in asynchronous interactions Evolutionary dynamics with game transitions.Proceedings of the National Academy of Sciences, 116:25398–25404, 2019
Reference 53
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation 0bfa9357-a221-4b80-aa85-629c6b5932ab · outbound
Direct reciprocity in asynchronous interactions Evolution of state-dependent strategies in stochastic games.Journal of Theoretical Biology, 527:110818, 2021
Reference 54
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation 36eab0f7-2a48-415e-9909-9e8cad2632ca · outbound
Direct reciprocity in asynchronous interactions Unresolved cited work
Reference 55
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation c4d16b95-0759-4224-ac13-11d453c300e4 · outbound
Direct reciprocity in asynchronous interactions The hydrogeomorphological effects of beaver dam-building activity
Reference 56
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation e28497db-73b3-4473-bc5f-122a398ee680 · outbound
Direct reciprocity in asynchronous interactions Elephants in the understory: opposing direct and indirect effects of consumption and ecosystem en- gineering by megaherbivores.Ecology, 97:3219–3230, 2016
Reference 57
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation 54e44df8-caad-400e-86b6-fceebca4e835 · outbound
Direct reciprocity in asynchronous interactions Evolution of cooperation in stochastic games.Nature, 559:246–249, 2018
Reference 58
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
Observation 271b768a-4a6b-4096-8f68-f4651e37b8a7 · outbound
Direct reciprocity in asynchronous interactions Multiple strategies in structured populations.Proceedings of the National Academy of Sciences, 108:2334–2337, 2011
Reference 59
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-07T06:34:17.273281+00:00.
No inbound Pith citation observations are available.