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Paper Citation Record · LEDGER

A non-reciprocal model for morphogenesis in symbiosis

As of 22 August 2026, this Paper Citation Record lists 38 of 38 outbound references and 0 inbound Pith citation observations for arXiv:2506.13299.

A citation records a reference. It does not transfer a finding from one paper to another.

pith.paper-citation-record.v1
2506.13299 v2

Coverage vector

measured 38 of 38 reference resolution

Typed states for the displayed outbound observations.

Source: paper_references, paper_reference_links, observed 2026-08-15T20:14:04.771216Z

measured 38 of 38 standing notices

One-hop event checks from named stored sources.

Source: scholarly_work_events, retraction_status_cache, observed 2026-08-22T06:32:14.747728+00:00

measured 0 of 0 inbound itemization

Pith citing papers itemized under the disclosed page cap.

Source: paper_references, paper_reference_links

measured 0 of 1 external citation measurements

A source-named dated measurement, never combined with another source.

Source: cited_works

Reference resolution

38 of 38 outbound references displayed

  • verified exact5
  • verified fuzzy16
  • unresolved16
  • parse uncertain0
  • malformed identifier1
  • metadata mismatch0

External citation measurements

No source-named external measurement is stored.

Outbound references

Observation 5ae859fb-a867-4e39-a933-70b7eacd7346 · outbound

This paper cites Nardon and H.

A non-reciprocal model for morphogenesis in symbiosis Nardon and H

Reference 1

Resolution
verified fuzzy
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Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.

source=pdf_text observed=2026-08-15T20:14:04.607998Z digest=sha256:83f0416d147395860d39025b218d8f2805f86a3219c8ce3001f7879e428a4ee6

Observation eea9c77d-975d-43db-88b4-111f4d9d7389 · outbound

This paper cites an unresolved cited work.

A non-reciprocal model for morphogenesis in symbiosis Unresolved cited work

Reference 2

Resolution
unresolved
raw_fallback, observed 2026-08-15T20:14:05.402895Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.

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Observation 52c7c0e9-d131-46aa-a308-1d5b70324249 · outbound

This paper cites Parniske, Arbuscular mycorrhiza: the mother of plant root endosymbioses, Nature Reviews Microbi- ology 6, 763 (2008), publisher: Nature Publishing Group.

A non-reciprocal model for morphogenesis in symbiosis Parniske, Arbuscular mycorrhiza: the mother of plant root endosymbioses, Nature Reviews Microbi- ology 6, 763 (2008), publisher: Nature Publishing Group

Reference 3

Resolution
verified fuzzy
raw_fallback, observed 2026-08-15T20:14:05.390185Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.

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Observation 9de10c7c-0f6d-40be-aa2e-3f6feec7ae89 · outbound

This paper cites an unresolved cited work.

A non-reciprocal model for morphogenesis in symbiosis Unresolved cited work

Reference 4

Resolution
unresolved
raw_fallback, observed 2026-08-15T20:14:05.378272Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.

source=pdf_text observed=2026-08-15T20:14:04.621946Z digest=sha256:f79df5d1bbee43758067fb6ae801cb818544265220d8411a715315e0a97f8d45

Observation ad86d61b-52a1-4238-8d19-59f48fc17cb0 · outbound

This paper cites an unresolved cited work.

A non-reciprocal model for morphogenesis in symbiosis Unresolved cited work

Reference 5

Resolution
unresolved
raw_fallback, observed 2026-08-15T20:14:05.366468Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.

source=pdf_text observed=2026-08-15T20:14:04.626238Z digest=sha256:1c1c4aebc9e1ea4b520574c47c6424d256d738270914661b585db16074fa6bc0

Observation afb1d1a3-d9ff-4636-aa02-aaeaee010c1c · outbound

This paper cites Margulis, Symbiosis and Evolution, Scientific American 225, 48 (1971), publisher: Scientific Amer- ican, a division of Nature America, Inc.

A non-reciprocal model for morphogenesis in symbiosis Margulis, Symbiosis and Evolution, Scientific American 225, 48 (1971), publisher: Scientific Amer- ican, a division of Nature America, Inc

Reference 6

Resolution
verified fuzzy
raw_fallback, observed 2026-08-15T20:14:05.354818Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.

source=pdf_text observed=2026-08-15T20:14:04.629952Z digest=sha256:e56135796fa2a9acd356d01c965db8d37cda8a92ea8ddc3d71920ce278baa3ce

Observation 250e0dd9-c328-446c-85f2-683ee91488cc · outbound

This paper cites an unresolved cited work.

A non-reciprocal model for morphogenesis in symbiosis Unresolved cited work

Reference 7

Resolution
unresolved
raw_fallback, observed 2026-08-15T20:14:05.343836Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.

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Observation b792fb40-bb27-4414-a7a7-944ffd8f7856 · outbound

This paper cites Fruchart, R.

A non-reciprocal model for morphogenesis in symbiosis Fruchart, R

Reference 8

Resolution
verified fuzzy
raw_fallback, observed 2026-08-15T20:14:05.331663Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.

source=pdf_text observed=2026-08-15T20:14:04.638447Z digest=sha256:22e5555b1554dedfe950371153ef4ac4a986a3bceabb0dffc18da02e3252449a

Observation 2270bdc9-cfae-40e3-b48b-387a9d95a895 · outbound

This paper cites an unresolved cited work.

A non-reciprocal model for morphogenesis in symbiosis Unresolved cited work

Reference 9

Resolution
unresolved
raw_fallback, observed 2026-08-15T20:14:05.321377Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.

source=pdf_text observed=2026-08-15T20:14:04.641971Z digest=sha256:42ea3cf9ae8259d831b0db34765f97a26b2d29c493f6a7fbf0e900fd4e10c5eb

Observation 9fcd063d-1eed-48de-9a16-3935b8c33930 · outbound

This paper cites Self-assembly of catalytically active colloidal molecules: Tailoring activity through surface chemistry.

A non-reciprocal model for morphogenesis in symbiosis Self-assembly of catalytically active colloidal molecules: Tailoring activity through surface chemistry

Reference 10

Resolution
verified exact
local_arxiv, observed 2026-08-15T20:14:05.036438Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.

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Observation 0b23328e-4c87-42b9-aa32-a6389f499f41 · outbound

This paper cites an unresolved cited work.

A non-reciprocal model for morphogenesis in symbiosis Unresolved cited work

Reference 11

Resolution
unresolved
raw_fallback, observed 2026-08-15T20:14:05.309301Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.

source=pdf_text observed=2026-08-15T20:14:04.649380Z digest=sha256:e84a9ad884a70274be9a2d7ad5e0412406a0333fc46202712c606b91f7ad06ed

Observation 51edd479-974d-4d53-be70-87e6a39928e0 · outbound

This paper cites an unresolved cited work.

A non-reciprocal model for morphogenesis in symbiosis Unresolved cited work

Reference 12

Resolution
unresolved
raw_fallback, observed 2026-08-15T20:14:05.295932Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.

source=pdf_text observed=2026-08-15T20:14:04.654129Z digest=sha256:be2ffd65a8e9d91de4b02747805fff8c8f0f42dc2b1d7a61efbb3a326a44871f

Observation a2a9548c-e633-4209-9510-13150f473582 · outbound

This paper cites Brauns and M.

A non-reciprocal model for morphogenesis in symbiosis Brauns and M

Reference 13

Resolution
verified fuzzy
raw_fallback, observed 2026-08-15T20:14:05.283184Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.

source=pdf_text observed=2026-08-15T20:14:04.658021Z digest=sha256:1de6d69631b72b2a3f50e3f89250c19e6aa6cc3d9f4ea30316ae9ffa8f6d0d4d

Observation 15d47c4b-72d5-4a12-8954-fa51a7acfff7 · outbound

This paper cites Osat and R.

A non-reciprocal model for morphogenesis in symbiosis Osat and R

Reference 14

Resolution
verified fuzzy
raw_fallback, observed 2026-08-15T20:14:05.270037Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.

source=pdf_text observed=2026-08-15T20:14:04.661869Z digest=sha256:3c744136f77a31e68e6e4f58f39234c3c8496f43aa798fb49112e02c165c501e

Observation 6f3d96a3-a785-46a9-bc7e-8ada19c09f4c · outbound

This paper cites Ouazan-Reboul, J.

A non-reciprocal model for morphogenesis in symbiosis Ouazan-Reboul, J

Reference 15

Resolution
verified fuzzy
raw_fallback, observed 2026-08-15T20:14:05.257246Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.

source=pdf_text observed=2026-08-15T20:14:04.666009Z digest=sha256:dd585d74aca16850d725e670802966a201e1bd4364ffc167de27a5bc9ad4c1aa

Observation 6cc0e2a9-d171-4054-8a24-76be3b3635dc · outbound

This paper cites an unresolved cited work.

A non-reciprocal model for morphogenesis in symbiosis Unresolved cited work

Reference 16

Resolution
unresolved
raw_fallback, observed 2026-08-15T20:14:05.246360Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.

source=pdf_text observed=2026-08-15T20:14:04.669640Z digest=sha256:24daade5efccdf41fff923c5d20da6da13c5d0a66ab13b49124807ff0d577180

Observation 11647d3a-d49c-49f0-a862-6e87f8434dec · outbound

This paper cites an unresolved cited work.

A non-reciprocal model for morphogenesis in symbiosis Unresolved cited work

Reference 17

Resolution
unresolved
raw_fallback, observed 2026-08-15T20:14:05.234983Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.

source=pdf_text observed=2026-08-15T20:14:04.673021Z digest=sha256:a56273bf7833fa7f503ac3cb6b3d11633c346e05948f1a3c0bce58effe8f31e0

Observation 9395ff7b-2cdb-4528-8e92-6421ba3c514b · outbound

This paper cites an unresolved cited work.

A non-reciprocal model for morphogenesis in symbiosis Unresolved cited work

Reference 18

Resolution
unresolved
raw_fallback, observed 2026-08-15T20:14:05.223558Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.

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Observation 2c4d7e9d-7ed1-4b21-b8d2-2e886d9f704b · outbound

This paper cites an unresolved cited work.

A non-reciprocal model for morphogenesis in symbiosis Unresolved cited work

Reference 19

Resolution
verified exact
doi, observed 2026-08-15T20:14:04.808025Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.

source=pdf_text observed=2026-08-15T20:14:04.681111Z digest=sha256:60e4e7ce20368ee1d2f6c49444098b1a4f59074b83ef5b3c8f063049f98a7285

Observation 4e70630f-f529-4a14-9ea9-221d4d1f8a93 · outbound

This paper cites Siggel, S.

A non-reciprocal model for morphogenesis in symbiosis Siggel, S

Reference 20

Resolution
verified fuzzy
raw_fallback, observed 2026-08-15T20:14:05.206942Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.

source=pdf_text observed=2026-08-15T20:14:04.685042Z digest=sha256:b82ca0ed498c148c7e909dfdf3b0861538e89ac48f527d25ed03e81fe720a050

Observation 6f35c023-7364-4738-954a-32a289531772 · outbound

This paper cites A tutorial for mesoscale computer simulations of lipid membranes: tether pulling, tubulation and fluctuation.

A non-reciprocal model for morphogenesis in symbiosis A tutorial for mesoscale computer simulations of lipid membranes: tether pulling, tubulation and fluctuation

Reference 21

Resolution
verified exact
local_arxiv, observed 2026-08-15T20:14:05.012812Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.

source=pdf_text observed=2026-08-15T20:14:04.690205Z digest=sha256:a8be4815d0c738fa16fae292f6adfb57e02f691995531e285b2d5d4a2f95e3f4

Observation ceef6504-fe4d-405a-ba8d-1a6a10eb4de3 · outbound

This paper cites Deserno, Elastic deformation of a fluid membrane upon colloid binding, Physical Review E69, 031903 (2004), publisher: American Physical Society.

A non-reciprocal model for morphogenesis in symbiosis Deserno, Elastic deformation of a fluid membrane upon colloid binding, Physical Review E69, 031903 (2004), publisher: American Physical Society

Reference 22

Resolution
verified fuzzy
raw_fallback, observed 2026-08-15T20:14:05.192465Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.

source=pdf_text observed=2026-08-15T20:14:04.696796Z digest=sha256:b0394bf97503ba769ea3047618dca1605fc03d3063a85fb48da0f940df5905fc

Observation 840b99c5-6be6-4812-89c7-b864e060c3fb · outbound

This paper cites Šarić and A.

A non-reciprocal model for morphogenesis in symbiosis Šarić and A

Reference 23

Resolution
verified fuzzy
raw_fallback, observed 2026-08-15T20:14:05.181043Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.

source=pdf_text observed=2026-08-15T20:14:04.701295Z digest=sha256:ed460670ddb24db0663e00f00fc266f7e7b5bbe3c2bbe1fcca3c84971b5c4aad

Observation fd439e3d-2589-4e62-b219-650ccafde9d0 · outbound

This paper cites Fluid membranes can drive linear aggregation of adsorbed spherical nanoparticles.

A non-reciprocal model for morphogenesis in symbiosis Fluid membranes can drive linear aggregation of adsorbed spherical nanoparticles

Reference 24

Resolution
verified exact
local_arxiv, observed 2026-08-15T20:14:04.982084Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.

source=pdf_text observed=2026-08-15T20:14:04.705205Z digest=sha256:44e3d1daa0dbbd7ee91304e62d5ab7ad99b68edefba572c8856d44cafdb1f3e3

Observation ad8fd08f-d9f3-4ebf-97d0-340784e50d2f · outbound

This paper cites an unresolved cited work.

A non-reciprocal model for morphogenesis in symbiosis Unresolved cited work

Reference 25

Resolution
unresolved
raw_fallback, observed 2026-08-15T20:14:05.170751Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.

source=pdf_text observed=2026-08-15T20:14:04.710628Z digest=sha256:6f3019e7b454c9d635a29a1c17257dbc3e505f386ee42c181d6c1918a2f14422

Observation 07fa6e16-6893-4ab6-b2bd-eb8594305240 · outbound

This paper cites an unresolved cited work.

A non-reciprocal model for morphogenesis in symbiosis Unresolved cited work

Reference 26

Resolution
unresolved
raw_fallback, observed 2026-08-15T20:14:05.160510Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.

source=pdf_text observed=2026-08-15T20:14:04.715020Z digest=sha256:47f620095bae08cd8dc2e4a246aa98112d78a654c7b604845a6478c2cb6393fb

Observation 4e9b0a81-5ad7-4845-82cd-4fea56aaefad · outbound

This paper cites Derényi, F.

A non-reciprocal model for morphogenesis in symbiosis Derényi, F

Reference 27

Resolution
verified fuzzy
raw_fallback, observed 2026-08-15T20:14:05.149941Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.

source=pdf_text observed=2026-08-15T20:14:04.719387Z digest=sha256:785614e515551aca09ae576cdb329f4d2d01f8bcd46dbebd8e98c4c480b7383f

Observation a08879cc-5066-4319-a043-d2d54b58da0a · outbound

This paper cites Rodrigues-Oliveira, F.

A non-reciprocal model for morphogenesis in symbiosis Rodrigues-Oliveira, F

Reference 28

Resolution
verified fuzzy
raw_fallback, observed 2026-08-15T20:14:05.138781Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.

source=pdf_text observed=2026-08-15T20:14:04.723958Z digest=sha256:fec0ece6da166870a8ec3ebe8e89dd4763166dbe336380d6b3736d3631c9e166

Observation f3b5336a-fdb0-48bc-a820-36efc43e0136 · outbound

This paper cites Muller, T.

A non-reciprocal model for morphogenesis in symbiosis Muller, T

Reference 29

Resolution
malformed identifier
no resolver link, observed 2026-08-15T20:14:04.728583Z

Source-reported events for the cited work

Unavailable: canonical work link unavailable.

source=pdf_text observed=2026-08-15T20:14:04.728583Z digest=sha256:7a48afe4ae87fffd025f07341ee7161d0f048675a2791ba0545ffd2ecb22b9b2

Observation 5709d15f-432b-498c-9885-0ae9097b5916 · outbound

This paper cites an unresolved cited work.

A non-reciprocal model for morphogenesis in symbiosis Unresolved cited work

Reference 30

Resolution
unresolved
raw_fallback, observed 2026-08-15T20:14:05.127883Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.

source=pdf_text observed=2026-08-15T20:14:04.732844Z digest=sha256:a97915141cb2ee30dd0e6800e68fff675ffd16d6273336198ba01601a96a5b4e

Observation f0911f5c-adad-4ec5-b22a-ebd754bcd9ed · outbound

This paper cites Hamada, K.

A non-reciprocal model for morphogenesis in symbiosis Hamada, K

Reference 31

Resolution
verified fuzzy
raw_fallback, observed 2026-08-15T20:14:05.116175Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.

source=pdf_text observed=2026-08-15T20:14:04.736727Z digest=sha256:ada0b865517e4a75c8acc5d773ab85a1ec9d29c9e689d4f37fc23c998955245a

Observation 5c86c3dc-0a45-419d-bd39-96938f20fae0 · outbound

This paper cites an unresolved cited work.

A non-reciprocal model for morphogenesis in symbiosis Unresolved cited work

Reference 32

Resolution
verified exact
raw_fallback, observed 2026-08-15T20:14:04.901155Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.

source=pdf_text observed=2026-08-15T20:14:04.740553Z digest=sha256:c554e75f311df32ee22f8628a1dbaad64bf171ec5d0632e176725f861d53bc08

Observation 48d2b8ba-6f7d-4d0e-ab30-dfe492cb86d1 · outbound

This paper cites an unresolved cited work.

A non-reciprocal model for morphogenesis in symbiosis Unresolved cited work

Reference 33

Resolution
unresolved
raw_fallback, observed 2026-08-15T20:14:05.105393Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.

source=pdf_text observed=2026-08-15T20:14:04.744731Z digest=sha256:ed256bba36ff1ff73f39acab8205617ffdc64c8f5087c18a78123b42c6207a01

Observation c15f3cde-3a96-451d-88de-f7f8e7ae9cd5 · outbound

This paper cites Imachi, M.

A non-reciprocal model for morphogenesis in symbiosis Imachi, M

Reference 34

Resolution
verified fuzzy
raw_fallback, observed 2026-08-15T20:14:05.094531Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.

source=pdf_text observed=2026-08-15T20:14:04.749144Z digest=sha256:2c1bd8b52923a26fb33a4ba04a798c2e66538b2d49a707b2a097f79248246800

Observation f52ad174-9267-401a-9420-9417b3ca1422 · outbound

This paper cites Spang, J.

A non-reciprocal model for morphogenesis in symbiosis Spang, J

Reference 35

Resolution
verified fuzzy
raw_fallback, observed 2026-08-15T20:14:05.083061Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.

source=pdf_text observed=2026-08-15T20:14:04.752996Z digest=sha256:2ef561eb57402f08abdb3c544b2dbc8b25bd550071d1a92e7093330f033640df

Observation 0c203619-a1d2-4d5c-9f2f-135028af2ad2 · outbound

This paper cites an unresolved cited work.

A non-reciprocal model for morphogenesis in symbiosis Unresolved cited work

Reference 36

Resolution
unresolved
raw_fallback, observed 2026-08-15T20:14:05.071976Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.

source=pdf_text observed=2026-08-15T20:14:04.761659Z digest=sha256:011153d1754cf320056b83f8a021c7d8136a9219dcac56e94dc6f502222cdad1

Observation 2b095e79-a4d4-4d18-b78b-be4a10ea5434 · outbound

This paper cites Wassermair, M.

A non-reciprocal model for morphogenesis in symbiosis Wassermair, M

Reference 37

Resolution
verified fuzzy
raw_fallback, observed 2026-08-15T20:14:05.060707Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.

source=pdf_text observed=2026-08-15T20:14:04.766675Z digest=sha256:c2ad4238f549b05fbbd7450ad14003b385783a536c42e076cd52690ceabf44e0

Observation 5d65578a-0de1-4eab-a67f-2d2fb20a4c94 · outbound

This paper cites an unresolved cited work.

A non-reciprocal model for morphogenesis in symbiosis Unresolved cited work

Reference 38

Resolution
unresolved
raw_fallback, observed 2026-08-15T20:14:05.047273Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.

source=pdf_text observed=2026-08-15T20:14:04.771216Z digest=sha256:8d5f02c7c5fcbabc9ef6cca2b2d1fd7f37a73f680594e8a343e690ac482ecfb4

Pith citing papers

No inbound Pith citation observations are available.