Typed states for the displayed outbound observations.
Source: paper_references, paper_reference_links, observed 2026-05-18T08:56:44.544404Z
Paper Citation Record · LEDGER
As of 6 August 2026, this Paper Citation Record lists 54 of 54 outbound references and 0 inbound Pith citation observations for arXiv:2510.07286.
A citation records a reference. It does not transfer a finding from one paper to another.
Typed states for the displayed outbound observations.
Source: paper_references, paper_reference_links, observed 2026-05-18T08:56:44.544404Z
One-hop event checks from named stored sources.
Source: scholarly_work_events, retraction_status_cache, observed 2026-08-06T06:34:29.942622+00:00
Pith citing papers itemized under the disclosed page cap.
Source: paper_references, paper_reference_links
A source-named dated measurement, never combined with another source.
Source: cited_works
54 of 54 outbound references displayed
External citation measurements
No source-named external measurement is stored.
Observation b424d2d2-c948-499c-9160-6e4d0d6177b9 · outbound
Evolutionary Profiles for Protein Fitness Prediction Correlated mutations and residue contacts in proteins.Proteins: Structure, Function, and Bioin- formatics, 18(4):309–317
Reference 1
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation d6bd861c-144a-4a62-9fcd-4293794c06da · outbound
Evolutionary Profiles for Protein Fitness Prediction Exploring protein fitness landscapes by directed evolution.Nature reviews Molecular cell biology, 10(12):866–876
Reference 2
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation 1781fb0e-d25f-4ac1-8a02-a02604c77564 · outbound
Evolutionary Profiles for Protein Fitness Prediction Machine learning for functional protein design.Nature biotechnology, 42(2):216–228
Reference 3
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation 11f3f0d5-ec41-4d57-8ce7-baebb772b370 · outbound
Evolutionary Profiles for Protein Fitness Prediction Low-n protein engineering with data-efficient deep learning.Nature methods, 18(4):389–396
Reference 4
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation 1211bcc1-4293-4df4-9392-5f1654347df7 · outbound
Evolutionary Profiles for Protein Fitness Prediction Mutation effects predicted from sequence co-variation.Nature biotechnology, 35(2):128–135
Reference 5
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation 09ea7c06-a8c4-4805-a528-aa960b357a13 · outbound
Evolutionary Profiles for Protein Fitness Prediction Language models enable zero-shot prediction of the effects of mutations on protein function.Advances in neural information processing systems, 34:29287–29303
Reference 6
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation 9fe71b03-d243-4798-949f-85cb17906abb · outbound
Evolutionary Profiles for Protein Fitness Prediction Multi-scale representation learning for protein fitness prediction
Reference 7
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation acff2c83-3604-4f19-93d2-824ecbce2eb4 · outbound
Evolutionary Profiles for Protein Fitness Prediction Lawrence Zitnick, Jerry Ma, and Rob Fergus
Reference 8
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation 34f18947-982d-4cba-8676-798129c8540e · outbound
Evolutionary Profiles for Protein Fitness Prediction Evolutionary-scale prediction of atomic level protein structure with a language model
Reference 9
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation 5b982290-edd9-4693-8e91-3acaab4ab98a · outbound
Evolutionary Profiles for Protein Fitness Prediction Learning inverse folding from millions of predicted structures
Reference 10
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation e5a5ccca-3495-4be7-a66e-06801b39b047 · outbound
Evolutionary Profiles for Protein Fitness Prediction Proteingym: 11 Large-scale benchmarks for protein fitness prediction and design.Advances in Neural Information Processing Systems, 36:64331–64379
Reference 11
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation 8d97c53e-1686-4446-9a1a-c1133fb5058f · outbound
Evolutionary Profiles for Protein Fitness Prediction Retrieval augmented protein language models for protein structure prediction
Reference 12
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation 66c0cc73-fc32-44e3-bb65-62bb9367522e · outbound
Evolutionary Profiles for Protein Fitness Prediction Retrieval-Enhanced Mutation Mastery: Augmenting Zero-Shot Prediction of Protein Language Model
Reference 13
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation eacfbdf7-c080-4c83-8a89-4bc2f986ecda · outbound
Evolutionary Profiles for Protein Fitness Prediction Multiple sequence alignment.Current Opinion in Structural Biology, 16(3):368–373
Reference 14
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation 9ec55ab6-82aa-45cd-a310-8704043b76aa · outbound
Evolutionary Profiles for Protein Fitness Prediction Esm-if1: Structure-informed protein language model for inverse folding
Reference 15
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation 20382257-1e2a-4519-8bee-9b0e10ecfeb3 · outbound
Evolutionary Profiles for Protein Fitness Prediction Algorithms for inverse reinforcement learning
Reference 16
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation baea7af9-a59a-48dc-98ee-612c02528796 · outbound
Evolutionary Profiles for Protein Fitness Prediction Ziebart, Andrew Maas, J
Reference 17
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation 915bb0eb-8699-40ef-829f-9bc872f7c97e · outbound
Evolutionary Profiles for Protein Fitness Prediction Fast and accurate protein structure search with foldseek.Nature biotechnology, 42(2):243–246
Reference 18
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation 0b9e1f73-52ea-416f-979f-a660a2bbf53c · outbound
Evolutionary Profiles for Protein Fitness Prediction R., Bruun, T
Reference 19
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation 78de7896-c85d-4f99-8866-ab82ba4c1358 · outbound
Evolutionary Profiles for Protein Fitness Prediction Advancing protein evolution with inverse folding models integrating structural and evolutionary constraints.Cell, 188(17):4674–4692.e19
Reference 20
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation ab0107bc-9288-4c28-9c2f-9996da217a94 · outbound
Evolutionary Profiles for Protein Fitness Prediction Deep mutational scanning: a new style of protein science.Nature Methods
Reference 21
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation ddcffeca-e861-4453-bec6-a77f241b0f92 · outbound
Evolutionary Profiles for Protein Fitness Prediction Semantical and geometrical protein encoding toward enhanced bioactivity and thermostability.Elife, 13:RP98033
Reference 22
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation 14b26bd8-a65a-49c1-989c-a05447a8fe60 · outbound
Evolutionary Profiles for Protein Fitness Prediction Saprot: Protein language modeling with structure-aware vocabulary.BioRxiv, pages 2023–10
Reference 23
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation cd2ed7e3-20d4-43e4-bcba-432dd7db75bf · outbound
Evolutionary Profiles for Protein Fitness Prediction ProSST: Protein language modeling with quantized structure and disentangled attention
Reference 24
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation e719a3be-ad8e-462c-815b-9965b5a6b5ab · outbound
Evolutionary Profiles for Protein Fitness Prediction Unresolved cited work
Reference 25
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation 3f3f612f-ce5d-48fb-8c55-3d73c57814ce · outbound
Evolutionary Profiles for Protein Fitness Prediction Diffusion language models are versatile protein learners
Reference 26
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation 0bd3e8a1-442f-44b2-91b0-9d5f048e8db3 · outbound
Evolutionary Profiles for Protein Fitness Prediction Language models enable zero-shot prediction of the effects of mutations on protein function
Reference 27
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation a3eaef52-1b05-404e-b9b9-efbb4791c057 · outbound
Evolutionary Profiles for Protein Fitness Prediction Epistasis in protein evolution.Protein science, 25(7):1204–1218
Reference 28
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation 223f83e9-4683-4014-97c6-60343a8d22b9 · outbound
Evolutionary Profiles for Protein Fitness Prediction Deep Think with Confidence
Reference 29
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation c0bb8668-c87c-40b2-aadf-56c56daf823f · outbound
Evolutionary Profiles for Protein Fitness Prediction Deep Researcher with Test-Time Diffusion
Reference 30
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation e8a351ff-d8bf-4700-b04a-c38a8979cdc8 · outbound
Evolutionary Profiles for Protein Fitness Prediction ParaThinker: Native Parallel Thinking as a New Paradigm to Scale LLM Test-time Compute
Reference 31
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation a6e26b5f-d64a-4902-a3dc-db33497432dc · outbound
Evolutionary Profiles for Protein Fitness Prediction The Majority is not always right: RL training for solution aggregation
Reference 32
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation f23c4df2-5c56-49cd-a9aa-b1958a937555 · outbound
Evolutionary Profiles for Protein Fitness Prediction Learning from Protein Structure with Geometric Vector Perceptrons
Reference 33
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation 1a4c5b36-3308-4867-91a8-c4029cb4e486 · outbound
Evolutionary Profiles for Protein Fitness Prediction Steering protein family design through profile bayesian flow
Reference 34
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation 26070dbb-ba38-4ff5-944e-5f1154be9070 · outbound
Evolutionary Profiles for Protein Fitness Prediction Unresolved cited work
Reference 35
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation e712a2e4-8307-4669-b48b-963a60d99545 · outbound
Evolutionary Profiles for Protein Fitness Prediction Qu, W., Ma, Y ., Ye, F., Lu, C., Zhou, Y ., Zhang, K., Wang, L., Gui, M., and Gu, Q
Reference 36
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation a70a2600-2adf-4453-b1c4-d3cd4dd15045 · outbound
Evolutionary Profiles for Protein Fitness Prediction AMix-1: A Pathway to Test-Time Scalable Protein Foundation Model
Reference 37
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation b67d4805-7f4b-44c2-adf1-850699311f61 · outbound
Evolutionary Profiles for Protein Fitness Prediction BERT: Pre-training of Deep Bidirectional Transformers for Language Understanding
Reference 38
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation e53949e5-37ec-40e3-a90c-6b87ecce7415 · outbound
Evolutionary Profiles for Protein Fitness Prediction Cath: increased structural coverage of functional space.Nucleic acids research, 49(D1): D266–D273
Reference 39
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation 9e05a98b-3028-4d0a-a717-273b66bb5c07 · outbound
Evolutionary Profiles for Protein Fitness Prediction Gemme: a simple and fast global epistatic model predicting mutational effects.Molecular biology and evolution, 36 (11):2604–2619
Reference 40
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation 382b7052-cba3-47c9-9771-aec65a478ffc · outbound
Evolutionary Profiles for Protein Fitness Prediction Progen2: exploring the boundaries of protein language models.Cell systems, 14(11):968–978
Reference 41
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation 25694cf7-95ba-4879-8291-48534370dff0 · outbound
Evolutionary Profiles for Protein Fitness Prediction Convolutions are competitive with transformers for protein sequence pretraining.Cell Systems, 15(3):286–294
Reference 42
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation af6a13ff-1661-45ce-b1b3-57db30866555 · outbound
Evolutionary Profiles for Protein Fitness Prediction Disease variant prediction with deep generative models of evolutionary data.Nature, 599(7883):91–95
Reference 43
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation f6048021-09a7-4e8d-95e9-80e68bad67fc · outbound
Evolutionary Profiles for Protein Fitness Prediction Msa transformer
Reference 44
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation 49c0f971-2f17-4799-830c-807a8cd4d5b6 · outbound
Evolutionary Profiles for Protein Fitness Prediction Tranception: protein fitness prediction with autoregressive transformers and inference-time retrieval
Reference 45
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation 3e534e18-7f72-40ca-babf-b7a138922cfc · outbound
Evolutionary Profiles for Protein Fitness Prediction Trancepteve: Combining family-specific and family-agnostic models of protein sequences for improved fitness prediction.bioRxiv, pages 2022–12
Reference 46
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation 2227a31d-866d-4c3e-90b0-8f7d4c9325bd · outbound
Evolutionary Profiles for Protein Fitness Prediction Robust deep learning–based protein sequence design using proteinmpnn.Science, 378 (6615):49–56
Reference 47
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation 66773503-748b-4232-8933-3329259f8521 · outbound
Evolutionary Profiles for Protein Fitness Prediction Masked inverse folding with sequence transfer for protein representation learning.Protein Engineering, Design and Selection, 36:gzad015
Reference 48
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation bb1068c0-2103-4bf5-bc16-e40528116cc3 · outbound
Evolutionary Profiles for Protein Fitness Prediction Deep generative models of genetic variation capture the effects of mutations.Nature methods, 15(10):816–822
Reference 49
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation 1cf29f73-86aa-4448-a883-1196cb6971eb · outbound
Evolutionary Profiles for Protein Fitness Prediction Evolutionary-scaleprediction of atomic-level protein structure with a language model.Science, 379(6637):1123–1130
Reference 50
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation b202a901-a872-4140-81f3-f3b9e9cc3410 · outbound
Evolutionary Profiles for Protein Fitness Prediction Muon is Scalable for LLM Training
Reference 51
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation 7ac52364-46eb-495e-a444-6c22c8898a72 · outbound
Evolutionary Profiles for Protein Fitness Prediction Decoupled Weight Decay Regularization
Reference 52
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation e1543698-35fa-4bbf-8e5a-415b4ab34763 · outbound
Evolutionary Profiles for Protein Fitness Prediction Alphafold protein structure database in 2024: providing structure coverage for over 214 million protein se- quences.Nucleic Acids Research, 52(D1):D368–D375
Reference 53
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
Observation 599b5fab-7ae3-460e-8539-7b34816a41dc · outbound
Evolutionary Profiles for Protein Fitness Prediction Unresolved cited work
Reference 54
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-06T06:34:29.942622+00:00.
No inbound Pith citation observations are available.