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Paper Citation Record · LEDGER

Probing 3D Chromatin Structure Awareness in Evo2 DNA Language Model

As of 12 August 2026, this Paper Citation Record lists 16 of 16 outbound references and 0 inbound Pith citation observations for arXiv:2604.07196.

A citation records a reference. It does not transfer a finding from one paper to another.

pith.paper-citation-record.v1
2604.07196 v1

Coverage vector

measured 16 of 16 reference resolution

Typed states for the displayed outbound observations.

Source: paper_references, paper_reference_links, observed 2026-05-10T17:55:29.224979Z

measured 16 of 16 standing notices

One-hop event checks from named stored sources.

Source: scholarly_work_events, retraction_status_cache, observed 2026-08-11T06:34:44.6726+00:00

measured 0 of 0 inbound itemization

Pith citing papers itemized under the disclosed page cap.

Source: paper_references, paper_reference_links

measured 0 of 1 external citation measurements

A source-named dated measurement, never combined with another source.

Source: cited_works

Reference resolution

16 of 16 outbound references displayed

  • verified exact0
  • verified fuzzy11
  • unresolved5
  • parse uncertain0
  • malformed identifier0
  • metadata mismatch0

External citation measurements

No source-named external measurement is stored.

Outbound references

Observation f9a67e8c-0e6c-40a0-b310-23ff23db62f6 · outbound

This paper cites and Cavalli, G.

Probing 3D Chromatin Structure Awareness in Evo2 DNA Language Model and Cavalli, G

Reference 1

Resolution
verified fuzzy
raw_fallback, observed 2026-05-17T08:14:18.084930Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-11T06:34:44.6726+00:00.

source=pdf_text observed=2026-05-10T17:55:29.224979Z digest=sha256:3c9ed5878edde6c2fe27a60d8266b940f1fde2220360fce57927c0172916197d

Observation 98bf7c2d-65bc-4429-8bba-ae7707665a61 · outbound

This paper cites P., Goodarzi, H., Hsu, P.

Probing 3D Chromatin Structure Awareness in Evo2 DNA Language Model P., Goodarzi, H., Hsu, P

Reference 2

Resolution
verified fuzzy
raw_fallback, observed 2026-05-17T08:14:18.063767Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-11T06:34:44.6726+00:00.

source=pdf_text observed=2026-05-10T17:55:29.224979Z digest=sha256:536a668fd43fd58d1663540e112f8120c438311b36da49de32906c47dcaa23a4

Observation 8a42e324-59c6-4e3b-a115-33110ebf2912 · outbound

This paper cites DNALONGBENCH: a bench- mark suite for long-range DNA prediction tasks.Nature Communications, 16(1):10108.

Probing 3D Chromatin Structure Awareness in Evo2 DNA Language Model DNALONGBENCH: a bench- mark suite for long-range DNA prediction tasks.Nature Communications, 16(1):10108

Reference 3

Resolution
verified fuzzy
raw_fallback, observed 2026-05-17T08:14:18.090871Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-11T06:34:44.6726+00:00.

source=pdf_text observed=2026-05-10T17:55:29.224979Z digest=sha256:2868924563782fc53de40aa996c799436dbc6806fbb3de72041fcd0270431475

Observation 1df5bac9-dcb9-4612-99de-0654adc4ca7a · outbound

This paper cites S., Meuleman, W., and Pinello, L.

Probing 3D Chromatin Structure Awareness in Evo2 DNA Language Model S., Meuleman, W., and Pinello, L

Reference 4

Resolution
verified fuzzy
raw_fallback, observed 2026-05-17T08:14:18.096667Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-11T06:34:44.6726+00:00.

source=pdf_text observed=2026-05-10T17:55:29.224979Z digest=sha256:410751c07d6b2b673099a18c2b019a28983430f793d5764ec4aa1f5c9aef4850

Observation e9876c3f-a23d-4ccd-9c00-3aa27419278b · outbound

This paper cites S., Stemmer-Rachamimov, A.

Probing 3D Chromatin Structure Awareness in Evo2 DNA Language Model S., Stemmer-Rachamimov, A

Reference 5

Resolution
verified fuzzy
raw_fallback, observed 2026-05-17T08:14:18.079015Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-11T06:34:44.6726+00:00.

source=pdf_text observed=2026-05-10T17:55:29.224979Z digest=sha256:36065d85575f1310b234c25f42ce69a383a24b98bead1262fc4428335e3f50ac

Observation 85764d58-d384-4f42-a009-bb5126b90a93 · outbound

This paper cites an unresolved cited work.

Probing 3D Chromatin Structure Awareness in Evo2 DNA Language Model Unresolved cited work

Reference 6

Resolution
unresolved
raw_fallback, observed 2026-05-17T08:14:18.107564Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-11T06:34:44.6726+00:00.

source=pdf_text observed=2026-05-10T17:55:29.224979Z digest=sha256:3b9c23d04fd27cad4ca26c01de2a9c05fc0308736034ca5d7211091bb35e0ed9

Observation fd1ed96e-7d23-4605-b77c-cc2762026161 · outbound

This paper cites an unresolved cited work.

Probing 3D Chromatin Structure Awareness in Evo2 DNA Language Model Unresolved cited work

Reference 7

Resolution
unresolved
raw_fallback, observed 2026-05-17T08:14:18.099581Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-11T06:34:44.6726+00:00.

source=pdf_text observed=2026-05-10T17:55:29.224979Z digest=sha256:296800ed8049e6fbfb43ca5a78ae50be5435d907c80aa20f3b6d8acf8fdb10b3

Observation 6b40512a-dfb8-46fc-ab49-b1f3917751fb · outbound

This paper cites an unresolved cited work.

Probing 3D Chromatin Structure Awareness in Evo2 DNA Language Model Unresolved cited work

Reference 8

Resolution
unresolved
raw_fallback, observed 2026-05-17T08:14:18.071026Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-11T06:34:44.6726+00:00.

source=pdf_text observed=2026-05-10T17:55:29.224979Z digest=sha256:8e9553d5fdd29582b03ca18cedda7fd685924ceb326eeeff6d1d1ecf67b25861

Observation 140c493e-12b5-40c6-ab25-850c8edfd61c · outbound

This paper cites CRISPR Inversion of CTCF Sites Alters Genome Topology and Enhancer/Promoter Function.Cell, 162(4):900–910.

Probing 3D Chromatin Structure Awareness in Evo2 DNA Language Model CRISPR Inversion of CTCF Sites Alters Genome Topology and Enhancer/Promoter Function.Cell, 162(4):900–910

Reference 9

Resolution
verified fuzzy
raw_fallback, observed 2026-05-17T08:14:18.073898Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-11T06:34:44.6726+00:00.

source=pdf_text observed=2026-05-10T17:55:29.224979Z digest=sha256:48d536d7df25d9c562af9c1a447c4679bebd40bed9f4261bcaf6b486cf83f04b

Observation b490e40d-7b22-4227-8143-a45ea46ff1f6 · outbound

This paper cites an unresolved cited work.

Probing 3D Chromatin Structure Awareness in Evo2 DNA Language Model Unresolved cited work

Reference 10

Resolution
unresolved
raw_fallback, observed 2026-05-17T08:14:18.076475Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-11T06:34:44.6726+00:00.

source=pdf_text observed=2026-05-10T17:55:29.224979Z digest=sha256:feb939907b1709896c37cfbd361a290245ca4f1a9de07e97c50594320a79f50e

Observation 5393c7b4-4d7a-4067-99d1-ec1255a194b1 · outbound

This paper cites S., Parsi, K.

Probing 3D Chromatin Structure Awareness in Evo2 DNA Language Model S., Parsi, K

Reference 11

Resolution
verified fuzzy
raw_fallback, observed 2026-05-17T08:14:18.102230Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-11T06:34:44.6726+00:00.

source=pdf_text observed=2026-05-10T17:55:29.224979Z digest=sha256:df80f2f110f39da0db6b70f21ade97da6704d3fcf93f943e1a5fb49538ebf2f8

Observation 9a0ac158-abfd-4a83-9eab-2e83a7cffe20 · outbound

This paper cites Post-mitotic transcriptional activation and 3D regulatory interactions show locus- and differentiation- specific sensitivity to cohesin depletion.bioRxiv.

Probing 3D Chromatin Structure Awareness in Evo2 DNA Language Model Post-mitotic transcriptional activation and 3D regulatory interactions show locus- and differentiation- specific sensitivity to cohesin depletion.bioRxiv

Reference 12

Resolution
verified fuzzy
raw_fallback, observed 2026-05-17T08:14:18.082192Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-11T06:34:44.6726+00:00.

source=pdf_text observed=2026-05-10T17:55:29.224979Z digest=sha256:84fb1d94deea02cf07b79db696a828de5d07ef593a63a0f906ff64fd943919a3

Observation c6fc6eee-e84f-4748-8441-e973c4f9010f · outbound

This paper cites A., Osterwalder, M., Franke, M., Timmermann, B., Hecht, J., Spielmann, M., Visel, A., and Mundlos, S.

Probing 3D Chromatin Structure Awareness in Evo2 DNA Language Model A., Osterwalder, M., Franke, M., Timmermann, B., Hecht, J., Spielmann, M., Visel, A., and Mundlos, S

Reference 13

Resolution
verified fuzzy
raw_fallback, observed 2026-05-17T08:14:18.105011Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-11T06:34:44.6726+00:00.

source=pdf_text observed=2026-05-10T17:55:29.224979Z digest=sha256:72030ccce254b3521abd961387359cd06372e7caec3092fd4bfd8f7a8970ce70

Observation ee6f1b20-244a-46ce-ba5d-dba34c0e3dfe · outbound

This paper cites Caduceus: Bi-Directional Equivariant Long- Range DNA Sequence Modeling.

Probing 3D Chromatin Structure Awareness in Evo2 DNA Language Model Caduceus: Bi-Directional Equivariant Long- Range DNA Sequence Modeling

Reference 14

Resolution
verified fuzzy
raw_fallback, observed 2026-05-17T08:14:18.087904Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-11T06:34:44.6726+00:00.

source=pdf_text observed=2026-05-10T17:55:29.224979Z digest=sha256:c1ddb047c438bea1de91f79244b27f7856a037b8552f4f42306de8544410bfe7

Observation 2cdccdb3-291b-480b-9d07-ed76a179fe20 · outbound

This paper cites an unresolved cited work.

Probing 3D Chromatin Structure Awareness in Evo2 DNA Language Model Unresolved cited work

Reference 15

Resolution
unresolved
raw_fallback, observed 2026-05-17T08:14:18.066886Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-11T06:34:44.6726+00:00.

source=pdf_text observed=2026-05-10T17:55:29.224979Z digest=sha256:07406ca190f1dfe67fbf3f089b3bf1fc6fed5bd5166cc7162c026e0ffddf0ed3

Observation a1ffde27-032b-4494-a995-4c0ef8a634fe · outbound

This paper cites Sequence-based modeling of three-dimensional genome architecture from kilobase to chromosome scale.

Probing 3D Chromatin Structure Awareness in Evo2 DNA Language Model Sequence-based modeling of three-dimensional genome architecture from kilobase to chromosome scale

Reference 16

Resolution
verified fuzzy
raw_fallback, observed 2026-05-17T08:14:18.093800Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-11T06:34:44.6726+00:00.

source=pdf_text observed=2026-05-10T17:55:29.224979Z digest=sha256:f9abaa9d3337f03434bf379a9e112355efe3f3bdd61776a656d0141e810d2380

Pith citing papers

No inbound Pith citation observations are available.