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Paper Citation Record · LEDGER

Simulating is not always understanding: When model complexity obscures biology

As of 22 August 2026, this Paper Citation Record lists 41 of 41 outbound references and 0 inbound Pith citation observations for arXiv:2608.06998.

A citation records a reference. It does not transfer a finding from one paper to another.

pith.paper-citation-record.v1
2608.06998 v1

Coverage vector

measured 41 of 41 reference resolution

Typed states for the displayed outbound observations.

Source: paper_references, paper_reference_links, observed 2026-08-10T16:57:58.841644Z

measured 41 of 41 standing notices

One-hop event checks from named stored sources.

Source: scholarly_work_events, retraction_status_cache, observed 2026-08-21T06:32:19.484+00:00

measured 0 of 0 inbound itemization

Pith citing papers itemized under the disclosed page cap.

Source: paper_references, paper_reference_links

measured 0 of 1 external citation measurements

A source-named dated measurement, never combined with another source.

Source: cited_works

Reference resolution

41 of 41 outbound references displayed

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External citation measurements

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Outbound references

Observation 46fa5d24-5148-43af-9471-87e601f83bb6 · outbound

This paper cites Dynamic mechanistic explanation: computational modeling of circadian rhythms as an exemplar for cognitive science.Studies in History and Philosophy of Science, 41:321–333, 2010.

Simulating is not always understanding: When model complexity obscures biology Dynamic mechanistic explanation: computational modeling of circadian rhythms as an exemplar for cognitive science.Studies in History and Philosophy of Science, 41:321–333, 2010

Reference 1

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Observation 8560b5fa-90a3-4ece-bb8e-701ef9ff07ce · outbound

This paper cites an unresolved cited work.

Simulating is not always understanding: When model complexity obscures biology Unresolved cited work

Reference 2

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Observation 20e8128a-2def-45f7-98e9-9634d1f86eca · outbound

This paper cites Cambridge University Press, Cambridge, 2nd edition, 2009.

Simulating is not always understanding: When model complexity obscures biology Cambridge University Press, Cambridge, 2nd edition, 2009

Reference 3

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Observation cffdfd75-c271-4e01-b6a9-496a8326413d · outbound

This paper cites Three kinds of idealization.Journal of Philosophy, 104:639–659, 2007.

Simulating is not always understanding: When model complexity obscures biology Three kinds of idealization.Journal of Philosophy, 104:639–659, 2007

Reference 4

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Observation ef705630-4e27-4c69-84d5-51ed4732b5fe · outbound

This paper cites Modelling and representing: an artefactual approach to model-based representation.

Simulating is not always understanding: When model complexity obscures biology Modelling and representing: an artefactual approach to model-based representation

Reference 5

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Observation ba20e472-a110-45ee-9a73-b6fdbf8caf04 · outbound

This paper cites Karr, Jayodita C.

Simulating is not always understanding: When model complexity obscures biology Karr, Jayodita C

Reference 6

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Observation 14ff04d9-41c0-4c9c-8e32-bfa6841ccca8 · outbound

This paper cites Thornburg, Ansel Maytin, Jonghan Kwon, Troy A.

Simulating is not always understanding: When model complexity obscures biology Thornburg, Ansel Maytin, Jonghan Kwon, Troy A

Reference 7

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Observation 025e50ac-d38a-431b-abf0-3bb7ce71619c · outbound

This paper cites Gutenkunst, Joshua J.

Simulating is not always understanding: When model complexity obscures biology Gutenkunst, Joshua J

Reference 8

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Observation 8fe705ac-e012-4a62-bb5f-a6343bec25d3 · outbound

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Simulating is not always understanding: When model complexity obscures biology Unresolved cited work

Reference 9

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Observation 07e9e287-3f67-4c3b-8cbc-685cfaa4af18 · outbound

This paper cites Adynamicalparadigmformolecularcellbiology.TrendsinCellBiology, 30:504–515, 2020.

Simulating is not always understanding: When model complexity obscures biology Adynamicalparadigmformolecularcellbiology.TrendsinCellBiology, 30:504–515, 2020

Reference 10

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Observation 59246984-fd78-47d0-b361-7ad160f97b47 · outbound

This paper cites Understanding from machine learning models.British Journal for the Philosophy of Science, 73(1):109–133, 2022.

Simulating is not always understanding: When model complexity obscures biology Understanding from machine learning models.British Journal for the Philosophy of Science, 73(1):109–133, 2022

Reference 11

Resolution
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Observation 0a9acca0-691f-4d2b-b101-d8f78348d36d · outbound

This paper cites Data-driven discovery of dynamical models in biology.Nature Reviews Physics, 2026.

Simulating is not always understanding: When model complexity obscures biology Data-driven discovery of dynamical models in biology.Nature Reviews Physics, 2026

Reference 12

Resolution
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Observation f5759892-4034-450c-8b73-7a03b66e34ba · outbound

This paper cites The strategy of model building in population biology.American Scientist, 54(4): 421–431, 1966.

Simulating is not always understanding: When model complexity obscures biology The strategy of model building in population biology.American Scientist, 54(4): 421–431, 1966

Reference 13

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Observation 4dbdf105-ece6-46b4-b684-a24abb0beb57 · outbound

This paper cites Nédélec, Thomas Surrey, Anthony C.

Simulating is not always understanding: When model complexity obscures biology Nédélec, Thomas Surrey, Anthony C

Reference 14

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Observation 3572f1b3-e5cf-4839-8516-0a3bc5b9f766 · outbound

This paper cites On the explanatory power of atomistic simulations.European Journal for Philosophy of Science, 15:45, 2025.

Simulating is not always understanding: When model complexity obscures biology On the explanatory power of atomistic simulations.European Journal for Philosophy of Science, 15:45, 2025

Reference 15

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Observation 1c345816-895e-4495-a645-6f990e2bb3ee · outbound

This paper cites Howtoinferexplanationsfromcomputersimulations.StudiesinHistoryandPhilosophy of Science Part A, 82:25–33, 2020.

Simulating is not always understanding: When model complexity obscures biology Howtoinferexplanationsfromcomputersimulations.StudiesinHistoryandPhilosophy of Science Part A, 82:25–33, 2020

Reference 16

Resolution
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Observation 8df609e4-12f4-4310-8fde-2fdc1b07eb2d · outbound

This paper cites Vertex models: from cell mechanics to tissue morphogenesis.Philosophical Transactions of the Royal Society B: Biological Sciences, 372(1720): 20150520, 2017.

Simulating is not always understanding: When model complexity obscures biology Vertex models: from cell mechanics to tissue morphogenesis.Philosophical Transactions of the Royal Society B: Biological Sciences, 372(1720): 20150520, 2017

Reference 17

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Observation ebd832ff-d0f0-425e-beda-2591aab4eb58 · outbound

This paper cites Cristina Marchetti, Jean-François Joanny, Sriram Ramaswamy, Tanniemola B.

Simulating is not always understanding: When model complexity obscures biology Cristina Marchetti, Jean-François Joanny, Sriram Ramaswamy, Tanniemola B

Reference 18

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Simulating is not always understanding: When model complexity obscures biology Unresolved cited work

Reference 19

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This paper cites Babtie and Michael P.

Simulating is not always understanding: When model complexity obscures biology Babtie and Michael P

Reference 20

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Simulating is not always understanding: When model complexity obscures biology Unresolved cited work

Reference 21

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Observation 983da84a-ec6b-47e1-8823-231a4aec3d91 · outbound

This paper cites The evolution of systems biology and systems medicine: from mechanistic models to uncertainty quantification.Annual Review of Biomedical Engineering, 27:425–447, 2025.

Simulating is not always understanding: When model complexity obscures biology The evolution of systems biology and systems medicine: from mechanistic models to uncertainty quantification.Annual Review of Biomedical Engineering, 27:425–447, 2025

Reference 22

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This paper cites Banga and Alejandro F.

Simulating is not always understanding: When model complexity obscures biology Banga and Alejandro F

Reference 23

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Observation 363649b4-19f7-4f9e-9b2e-ad883b37937c · outbound

This paper cites A modular approach for modeling the cell cycle based on functional response curves.PLoS Computational Biology, 17:e1009008, 2021.

Simulating is not always understanding: When model complexity obscures biology A modular approach for modeling the cell cycle based on functional response curves.PLoS Computational Biology, 17:e1009008, 2021

Reference 24

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Observation 5a9ebda1-62a4-432f-9fb1-15b3a5b98eb9 · outbound

This paper cites Bistable,biphasicregulationofPP2A-B55accounts for the dynamics of mitotic substrate phosphorylation.Current Biology, 31(4):794–808.e6, 2021.

Simulating is not always understanding: When model complexity obscures biology Bistable,biphasicregulationofPP2A-B55accounts for the dynamics of mitotic substrate phosphorylation.Current Biology, 31(4):794–808.e6, 2021

Reference 25

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This paper cites Simulation and the sense of understanding.

Simulating is not always understanding: When model complexity obscures biology Simulation and the sense of understanding

Reference 26

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Observation b15c7098-424c-4ad7-b560-bd1617bd7701 · outbound

This paper cites No understanding without explanation.Studies in History and Philosophy of Science, 44:510–515, 2013.

Simulating is not always understanding: When model complexity obscures biology No understanding without explanation.Studies in History and Philosophy of Science, 44:510–515, 2013

Reference 27

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Observation 0d785b0d-23fa-4750-bc2e-a690d82d4aad · outbound

This paper cites de Regt.Understanding Scientific Understanding.

Simulating is not always understanding: When model complexity obscures biology de Regt.Understanding Scientific Understanding

Reference 28

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verified fuzzy
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Observation 62c07d43-ba81-4c0d-ba87-def785e4b793 · outbound

This paper cites Batterman and Collin C.

Simulating is not always understanding: When model complexity obscures biology Batterman and Collin C

Reference 29

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verified fuzzy
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No event found in the named queried sources as of 2026-08-21T06:32:19.484+00:00.

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Observation c4759552-fead-4890-8d75-a43bc49fcf44 · outbound

This paper cites Complex versus complicated systems biology, universality versus detailed modeling.

Simulating is not always understanding: When model complexity obscures biology Complex versus complicated systems biology, universality versus detailed modeling

Reference 30

Resolution
verified fuzzy
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No event found in the named queried sources as of 2026-08-21T06:32:19.484+00:00.

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Observation f63cb80b-cadc-45f7-b3c6-59c4a49f62fc · outbound

This paper cites Craver and David M.

Simulating is not always understanding: When model complexity obscures biology Craver and David M

Reference 31

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Observation 94c9ac0b-0a77-4016-9d9e-863254c5028f · outbound

This paper cites Beyond reduction and pluralism: toward an epistemology of explanatory integration in biology.Erkenntnis, 73:295–311, 2010.

Simulating is not always understanding: When model complexity obscures biology Beyond reduction and pluralism: toward an epistemology of explanatory integration in biology.Erkenntnis, 73:295–311, 2010

Reference 32

Resolution
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No event found in the named queried sources as of 2026-08-21T06:32:19.484+00:00.

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Observation 5bc8d2d1-47dc-4498-ba7c-af8134815169 · outbound

This paper cites Theinfluence of cell mechanics, cell–cell interactions, and proliferation on epithelial packing.Current Biology, 17: 2095–2104, 2007.

Simulating is not always understanding: When model complexity obscures biology Theinfluence of cell mechanics, cell–cell interactions, and proliferation on epithelial packing.Current Biology, 17: 2095–2104, 2007

Reference 33

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Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-21T06:32:19.484+00:00.

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Observation d0ff1c0d-ca1f-471c-91e6-2f354f0c179b · outbound

This paper cites Burkhardt, et al.

Simulating is not always understanding: When model complexity obscures biology Burkhardt, et al

Reference 34

Resolution
verified fuzzy
raw_fallback, observed 2026-08-10T16:57:58.994359Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-21T06:32:19.484+00:00.

source=pdf_text observed=2026-08-10T16:57:58.809412Z digest=sha256:266535b1976299bf349031d062d069b1848aa0548df92d9464bb50cda617c567

Observation 8e32d83a-6100-4f6d-a0da-c0b561466789 · outbound

This paper cites scgpt: toward building a foundation model for single-cell multi-omics using generative AI.Nature Methods, 21:1470–1480, 2024.

Simulating is not always understanding: When model complexity obscures biology scgpt: toward building a foundation model for single-cell multi-omics using generative AI.Nature Methods, 21:1470–1480, 2024

Reference 35

Resolution
verified fuzzy
raw_fallback, observed 2026-08-10T16:57:58.980477Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-21T06:32:19.484+00:00.

source=pdf_text observed=2026-08-10T16:57:58.814107Z digest=sha256:e2097cf6a94de6f921103280160eff23be4d6ad31e9a9c36d62db514b859fbed

Observation d992ef03-1685-4a12-b61e-ba64f48463c5 · outbound

This paper cites Brydon, Zexian Zeng, X.

Simulating is not always understanding: When model complexity obscures biology Brydon, Zexian Zeng, X

Reference 36

Resolution
verified fuzzy
raw_fallback, observed 2026-08-10T16:57:58.964332Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-21T06:32:19.484+00:00.

source=pdf_text observed=2026-08-10T16:57:58.818805Z digest=sha256:bc0166814a644cc5dd4571a71d4afb1eaf426ce567b08c9ad7b76a51855609e7

Observation a19e57c4-6d14-4de8-9874-512373f1bb21 · outbound

This paper cites Towards the construction of a virtual yeast.Nature, 655: 59–70, 2026.

Simulating is not always understanding: When model complexity obscures biology Towards the construction of a virtual yeast.Nature, 655: 59–70, 2026

Reference 37

Resolution
verified fuzzy
raw_fallback, observed 2026-08-10T16:57:58.947270Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-21T06:32:19.484+00:00.

source=pdf_text observed=2026-08-10T16:57:58.823284Z digest=sha256:f8405099ee42187e6c53fa9cc3488f8532a24a1481b693da38ba67d52de096db

Observation 06bd4e12-3c16-4ef9-85cd-3848ac2bd51e · outbound

This paper cites Garcia Gomez, Kirsten H.

Simulating is not always understanding: When model complexity obscures biology Garcia Gomez, Kirsten H

Reference 38

Resolution
verified fuzzy
raw_fallback, observed 2026-08-10T16:57:58.931108Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-21T06:32:19.484+00:00.

source=pdf_text observed=2026-08-10T16:57:58.827862Z digest=sha256:11bf33dc76c811a0b4e735c9e3340012160420f562ffc9fe55b609d958215a0b

Observation 0f7b561c-4751-4600-a1e1-693b7076faa0 · outbound

This paper cites Artificial intelligence-enabled multi-scale virtual cell: perspective, challenges, and opportunities.Briefings in Bioinformatics, 27(bbag104), 2026.

Simulating is not always understanding: When model complexity obscures biology Artificial intelligence-enabled multi-scale virtual cell: perspective, challenges, and opportunities.Briefings in Bioinformatics, 27(bbag104), 2026

Reference 39

Resolution
verified fuzzy
raw_fallback, observed 2026-08-10T16:57:58.914340Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-21T06:32:19.484+00:00.

source=pdf_text observed=2026-08-10T16:57:58.832266Z digest=sha256:4e26ee57da264dac196bf153a11700f8b7d01cd5947b12513ce26e322499ce60

Observation dcf7750a-f1ed-4c46-a376-73ecacd286ae · outbound

This paper cites Theis, and Bo Wang.

Simulating is not always understanding: When model complexity obscures biology Theis, and Bo Wang

Reference 40

Resolution
verified fuzzy
raw_fallback, observed 2026-08-10T16:57:58.898410Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-21T06:32:19.484+00:00.

source=pdf_text observed=2026-08-10T16:57:58.837162Z digest=sha256:732a7fdfd937c56b5c5b49d15540be6bcaa7d6163322b5bb2df913888737e371

Observation 6367b7b2-7eb0-44e2-ad4f-cd9856a4c6d5 · outbound

This paper cites Gitlab repository.https://gitlab.kuleuven.be/gelenslab/publications/ 2026_Gelens_perspective_simulating_and_understanding.git, 2026.

Simulating is not always understanding: When model complexity obscures biology Gitlab repository.https://gitlab.kuleuven.be/gelenslab/publications/ 2026_Gelens_perspective_simulating_and_understanding.git, 2026

Reference 41

Resolution
verified fuzzy
raw_fallback, observed 2026-08-10T16:57:58.881999Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-21T06:32:19.484+00:00.

source=pdf_text observed=2026-08-10T16:57:58.841644Z digest=sha256:afc2f37273f62555b0e7f65f70feefc145e37853dc01bae482a0c4ddccff1fc1

Pith citing papers

No inbound Pith citation observations are available.