Paper Citation Record · LEDGER

Self-propulsive active nematics

As of 13 August 2026, this Paper Citation Record lists 45 of 45 outbound references and 0 inbound Pith citation observations for arXiv:2509.02386.

A citation records a reference. It does not transfer a finding from one paper to another.

pith.paper-citation-record.v1
2509.02386 v1

Coverage vector

measured 45 of 45 reference resolution

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Source: paper_references, paper_reference_links, observed 2026-08-05T11:47:37.995870Z

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Pith citing papers itemized under the disclosed page cap.

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Reference resolution

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Outbound references

Observation 810197bc-03c6-49d8-9c70-ac7b7377e97a · outbound

This paper cites 2017 Active matter at the interface between materials science and cell biology.Nature Reviews Materials2, 1–14.

Self-propulsive active nematics 2017 Active matter at the interface between materials science and cell biology.Nature Reviews Materials2, 1–14

Reference 1

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Observation 69765497-785d-4fd3-b998-46a280d88449 · outbound

This paper cites giant number fluctuations.

Self-propulsive active nematics giant number fluctuations

Reference 2

Resolution: verified fuzzy; raw_fallback, observed 2026-08-05T11:47:38.707008Z

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Observation 63a150d2-67d8-4dbb-9adf-7238ed7bae92 · outbound

This paper cites 2014 Living liquid crystals.Proceed- ings of the National Academy of Sciences111, 1265–1270.

Self-propulsive active nematics 2014 Living liquid crystals.Proceed- ings of the National Academy of Sciences111, 1265–1270

Reference 3

Resolution: verified exact; doi, observed 2026-08-05T11:47:38.576653Z

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Observation f34e04fe-a360-4cca-8055-b17a70b4b241 · outbound

This paper cites 2022 Bacterial active matter.Reports on Progress in Physics85, 076601.

Self-propulsive active nematics 2022 Bacterial active matter.Reports on Progress in Physics85, 076601

Reference 4

Resolution: verified exact; doi, observed 2026-08-05T11:47:38.591845Z

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Observation 78a11e96-ee84-4a1e-b612-dca255201a62 · outbound

This paper cites 2020 Dense active matter model of motion patterns in confluent cell monolayers.Nature Communications 11, 1405.

Self-propulsive active nematics 2020 Dense active matter model of motion patterns in confluent cell monolayers.Nature Communications 11, 1405

Reference 5

Resolution: unresolved; no resolver link, observed 2026-08-05T11:47:37.840827Z

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Observation e077bde8-caa1-471c-bae5-e5426ffa65ab · outbound

This paper cites 2018 Turbulent Dynamics of Epithelial Cell Cultures.Physical Review Letters120.

Self-propulsive active nematics 2018 Turbulent Dynamics of Epithelial Cell Cultures.Physical Review Letters120

Reference 6

Resolution: unresolved; no resolver link, observed 2026-08-05T11:47:37.835929Z

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Observation 5c7633b3-9b8c-4e77-a0b4-455b10b62580 · outbound

This paper cites 2016 Emergent behavior in active colloids.Journal of Physics: Con- densed Matter28, 253001.

Self-propulsive active nematics 2016 Emergent behavior in active colloids.Journal of Physics: Con- densed Matter28, 253001

Reference 7

Resolution: unresolved; no resolver link, observed 2026-08-05T11:47:37.849609Z

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Observation 4c4b6dd7-b033-4f74-9532-6217581f0aa3 · outbound

This paper cites 2020 The 2020 motile active matter roadmap.

Self-propulsive active nematics 2020 The 2020 motile active matter roadmap

Reference 8

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Observation a37d3b94-c014-402c-9815-179f6218e37f · outbound

This paper cites 2022 Physics of liquid crystals in cell biology.Trends in Cell Biology32, 140–150.

Self-propulsive active nematics 2022 Physics of liquid crystals in cell biology.Trends in Cell Biology32, 140–150

Reference 9

Resolution: unresolved; no resolver link, observed 2026-08-05T11:47:37.857619Z

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Observation e4814c04-ff10-406d-9e30-9b23c741cf6c · outbound

This paper cites 2018 Biological tissues as active nematic liquid crystals.

Self-propulsive active nematics 2018 Biological tissues as active nematic liquid crystals

Reference 10

Resolution: verified exact; doi, observed 2026-08-05T11:47:38.522726Z

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Observation e298540e-7639-4d5c-9be7-41405ca73360 · outbound

This paper cites 2014 Defect dy- namics in active nematics.Philosophical Transactions of the Royal Society A: Mathe- matical, Physical and Engineering Sciences372, 20130365.

Self-propulsive active nematics 2014 Defect dy- namics in active nematics.Philosophical Transactions of the Royal Society A: Mathe- matical, Physical and Engineering Sciences372, 20130365

Reference 11

Resolution: verified exact; raw_fallback, observed 2026-08-05T11:47:38.692229Z

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Observation 76c88ce4-bb40-461e-90e8-1aa5d4bb7444 · outbound

This paper cites 2019 Emergence of active nematics in chaining bacterial biofilms.

Self-propulsive active nematics 2019 Emergence of active nematics in chaining bacterial biofilms

Reference 12

Resolution: malformed identifier; no resolver link, observed 2026-08-05T11:47:37.861632Z

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Observation 28f1a76e-88ac-41ec-9483-8d3935ccb618 · outbound

This paper cites 2018 Active nematics.

Self-propulsive active nematics 2018 Active nematics

Reference 13

Resolution: unresolved; no resolver link, observed 2026-08-05T11:47:37.873971Z

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Observation 752bbd39-38d2-4519-b423-515bd7bf1771 · outbound

This paper cites 2022 Active Turbulence.Annual Review of Con- densed Matter Physics13, 143–170.

Self-propulsive active nematics 2022 Active Turbulence.Annual Review of Con- densed Matter Physics13, 143–170

Reference 14

Resolution: verified exact; doi, observed 2026-08-05T11:47:38.489604Z

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Observation 2c41e155-f525-4bb8-9c38-53867096c37c · outbound

This paper cites 2021 Bac- teria solve the problem of crowding by moving slowly.Nature Physics 17, 205–210.

Self-propulsive active nematics 2021 Bac- teria solve the problem of crowding by moving slowly.Nature Physics 17, 205–210

Reference 15

Resolution: unresolved; no resolver link, observed 2026-08-05T11:47:37.882719Z

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Observation 0d189b58-c66b-45d3-8c33-a001af472638 · outbound

This paper cites 2024 Topological de- fects lead to energy transfer in active nematics.

Self-propulsive active nematics 2024 Topological de- fects lead to energy transfer in active nematics

Reference 16

Resolution: verified exact; doi, observed 2026-08-05T11:47:38.464845Z

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Observation 8d72b38f-18b1-4a97-9727-c02484f77ab3 · outbound

This paper cites 2018 Mesoscale physical principles of collective cell organization.

Self-propulsive active nematics 2018 Mesoscale physical principles of collective cell organization

Reference 17

Resolution: unresolved; no resolver link, observed 2026-08-05T11:47:37.890951Z

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Observation 7854275b-1ed6-4e1f-baa8-585877a51473 · outbound

This paper cites 2021 Topological defects pro- mote layer formation in Myxococcus xanthus colonies.Nature Physics 17, 211–215.

Self-propulsive active nematics 2021 Topological defects pro- mote layer formation in Myxococcus xanthus colonies.Nature Physics 17, 211–215

Reference 18

Resolution: verified exact; doi, observed 2026-08-05T11:47:38.375689Z

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Observation b9e39271-fe09-4510-9ea5-8528598aa8a5 · outbound

This paper cites 2015 Turbulence of swarm- ing sperm.Physical Revew E92, 032722.

Self-propulsive active nematics 2015 Turbulence of swarm- ing sperm.Physical Revew E92, 032722

Reference 19

Resolution: verified exact; doi, observed 2026-08-05T11:47:38.339188Z

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Observation 32b85d55-fe7c-4413-a72f-b1f722708fb3 · outbound

This paper cites 2017 Topological defects control collective dy- namics in neural progenitor cell cultures.Nature 545, 327–331.

Self-propulsive active nematics 2017 Topological defects control collective dy- namics in neural progenitor cell cultures.Nature 545, 327–331

Reference 20

Resolution: unresolved; no resolver link, observed 2026-08-05T11:47:37.895152Z

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Observation d2fd787f-2fbf-4aed-9760-c26499fc3d66 · outbound

This paper cites 2012 Self-regulation in self-propelled nematic fluids.The European Physical Journal E35, 1–8.

Self-propulsive active nematics 2012 Self-regulation in self-propelled nematic fluids.The European Physical Journal E35, 1–8

Reference 21

Resolution: verified exact; doi, observed 2026-08-05T11:47:38.314293Z

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Observation 1489f28a-572c-4189-8b3f-8daaebf9c85d · outbound

This paper cites 2008 Hydrodynamics of self-propelled hard rods.Physical Review E—Statistical, Nonlinear, and Soft Matter Physics77, 011920.

Self-propulsive active nematics 2008 Hydrodynamics of self-propelled hard rods.Physical Review E—Statistical, Nonlinear, and Soft Matter Physics77, 011920

Reference 22

Resolution: malformed identifier; no resolver link, observed 2026-08-05T11:47:37.903150Z

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Observation 08e064e5-5617-4b5e-892b-fb0b42e50185 · outbound

This paper cites Phase diagram, confining strings, and a new universality class in nematopolar matter.

Self-propulsive active nematics Phase diagram, confining strings, and a new universality class in nematopolar matter

Reference 23

Resolution: verified exact; local_arxiv, observed 2026-08-05T11:47:38.285035Z

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No event found in the named queried sources as of 2026-08-13T06:32:02.005865+00:00.

Observation e06dcc23-a2c2-4df5-a930-fabad00b93b5 · outbound

This paper cites 2020 Self-Propelled Rods: Insights and Perspectives for Active Matter.Annual Review of Condensed Matter Physics11, 441–466.

Self-propulsive active nematics 2020 Self-Propelled Rods: Insights and Perspectives for Active Matter.Annual Review of Condensed Matter Physics11, 441–466

Reference 24

Resolution: verified exact; doi, observed 2026-08-05T11:47:38.299881Z

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Observation ad96ff80-92b8-4656-ba48-d56bd2c4b6a5 · outbound

This paper cites 2012 Polar patterns in active fluids.Soft Matter8, 129–139.

Self-propulsive active nematics 2012 Polar patterns in active fluids.Soft Matter8, 129–139

Reference 25

Resolution: verified exact; doi, observed 2026-08-05T11:47:38.255698Z

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Observation 9e536e5c-8eec-44ac-aafb-367b8e3c75cb · outbound

This paper cites 2022 Unifying polar and nematic active matter: emergence and co-existence of half-integer and full-integer topological de- fects.

Self-propulsive active nematics 2022 Unifying polar and nematic active matter: emergence and co-existence of half-integer and full-integer topological de- fects

Reference 26

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Observation 1d877c86-4c7f-4707-8a31-7dbc994761a6 · outbound

This paper cites This behaviour is also shared for bacterium, such asMyxococcus xanthus, where each individual particle aligns with the flow direction [29].

Self-propulsive active nematics This behaviour is also shared for bacterium, such asMyxococcus xanthus, where each individual particle aligns with the flow direction [29]

Reference 27

Resolution: verified fuzzy; raw_fallback, observed 2026-08-05T11:47:38.721601Z

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Observation b79eaae4-5743-41db-abaf-0d3d4e180d17 · outbound

This paper cites 2013 Hydrodynamics of soft active matter.Reviews of Modern Physics85, 1143–1189.

Self-propulsive active nematics 2013 Hydrodynamics of soft active matter.Reviews of Modern Physics85, 1143–1189

Reference 28

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Observation 5e35d854-3288-4e69-9a16-f08508d5f51a · outbound

This paper cites 2003 Shear flow-induced motility of Dictyostelium discoideum cells on solid substrate.Journal of Cell Science 116, 4331–4338.

Self-propulsive active nematics 2003 Shear flow-induced motility of Dictyostelium discoideum cells on solid substrate.Journal of Cell Science 116, 4331–4338

Reference 29

Resolution: verified exact; doi, observed 2026-08-05T11:47:38.231053Z

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Observation 768abd3e-097e-4bf7-b820-0a3fd6c5f191 · outbound

This paper cites 2019 Sustained oscillations of epithelial cell sheets.

Self-propulsive active nematics 2019 Sustained oscillations of epithelial cell sheets

Reference 30

Resolution: unresolved; no resolver link, observed 2026-08-05T11:47:37.936258Z

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Observation c0894144-bb5d-4645-82c4-017b8df855d3 · outbound

This paper cites Local polar order controls mechanical stress and triggers layer formation in developing Myxococcus xanthus colonies.

Self-propulsive active nematics Local polar order controls mechanical stress and triggers layer formation in developing Myxococcus xanthus colonies

Reference 31

Resolution: verified exact; local_arxiv, observed 2026-08-05T11:47:38.206161Z

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Observation 6a69401c-fd2a-4590-a778-96a73f6b721f · outbound

This paper cites (10.1103/PhysRevE.76.031921).

Self-propulsive active nematics (10.1103/PhysRevE.76.031921)

Reference 32

Resolution: verified exact; doi, observed 2026-08-05T11:47:38.185582Z

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Observation 9f06a998-768f-4ed9-a9f5-28deb403ddfe · outbound

This paper cites 2012 Banding, excitability and chaos in active nematic suspensions.

Self-propulsive active nematics 2012 Banding, excitability and chaos in active nematic suspensions

Reference 33

Resolution: malformed identifier; no resolver link, observed 2026-08-05T11:47:37.948479Z

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Observation e79b3ffc-c90b-4b24-95de-dc97fa6e7929 · outbound

This paper cites 2020 Activity Induced Nematic Order in Isotropic Liquid Crystals.

Self-propulsive active nematics 2020 Activity Induced Nematic Order in Isotropic Liquid Crystals

Reference 34

Resolution: verified exact; doi, observed 2026-08-05T11:47:38.161768Z

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Observation 85eb0f97-20fa-45f4-ba28-898af0e383d4 · outbound

This paper cites 2021 Inertia Drives a Flocking Phase Transition in Viscous Active Fluids.Physical Review X 11, 031063.

Self-propulsive active nematics 2021 Inertia Drives a Flocking Phase Transition in Viscous Active Fluids.Physical Review X 11, 031063

Reference 35

Resolution: verified exact; doi, observed 2026-08-05T11:47:38.148018Z

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Observation ea3a6cb3-2a32-4f8c-a12d-5cdef6857bd4 · outbound

This paper cites 2015 Intrinsic free energy in active nematics.

Self-propulsive active nematics 2015 Intrinsic free energy in active nematics

Reference 36

Resolution: malformed identifier; no resolver link, observed 2026-08-05T11:47:37.960613Z

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Observation e623819b-dc49-44c7-8b88-650e750abd9e · outbound

This paper cites 2018 Hyperuniform states of matter.

Self-propulsive active nematics 2018 Hyperuniform states of matter

Reference 37

Resolution: unresolved; no resolver link, observed 2026-08-05T11:47:37.964534Z

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Observation 941d195e-b477-48c4-82dd-c670aa7b59c1 · outbound

This paper cites 2021 Local Number Fluctuations in Hyperuniform and Nonhyperuniform Systems: Higher-Order Moments and Distribution Functions.

Self-propulsive active nematics 2021 Local Number Fluctuations in Hyperuniform and Nonhyperuniform Systems: Higher-Order Moments and Distribution Functions

Reference 38

Resolution: verified exact; doi, observed 2026-08-05T11:47:38.114470Z

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No event found in the named queried sources as of 2026-08-13T06:32:02.005865+00:00.

Observation 97fbc8fc-a873-416d-a3dc-95bf92ed6a26 · outbound

This paper cites 1995 Long-Range Order in a Two-Dimensional DynamicalXY Model: How Birds Fly Together.Physical Review Letters75, 4326–4329.

Self-propulsive active nematics 1995 Long-Range Order in a Two-Dimensional DynamicalXY Model: How Birds Fly Together.Physical Review Letters75, 4326–4329

Reference 39

Resolution: unresolved; no resolver link, observed 2026-08-05T11:47:37.972579Z

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Observation dba60c90-5a4a-42c7-b1c9-d47d09ce6e99 · outbound

This paper cites 2015 Geometry and topology of turbulence in active nematics.Physical Re- view X 5, 031003.

Self-propulsive active nematics 2015 Geometry and topology of turbulence in active nematics.Physical Re- view X 5, 031003

Reference 40

Resolution: unresolved; no resolver link, observed 2026-08-05T11:47:37.976593Z

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Observation 7e441b3d-a39c-42d7-850a-e4be5befa8eb · outbound

This paper cites 2015 New class of turbulence in active fluids.Proceedings of the National Academy of Sciences112, 15048–15053.

Self-propulsive active nematics 2015 New class of turbulence in active fluids.Proceedings of the National Academy of Sciences112, 15048–15053

Reference 41

Resolution: unresolved; no resolver link, observed 2026-08-05T11:47:37.980417Z

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Observation 1693bb3a-b562-4d87-adc1-c3c584d5921c · outbound

This paper cites 2017 Topological defects in epithelia govern cell death and extrusion.Nature 544, 212–216.

Self-propulsive active nematics 2017 Topological defects in epithelia govern cell death and extrusion.Nature 544, 212–216

Reference 42

Resolution: unresolved; no resolver link, observed 2026-08-05T11:47:37.984272Z

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Observation 79bf3abf-bc80-402d-a941-9998fa6ace61 · outbound

This paper cites 2021 Cell migration guided by long-lived spatial memory.Nature Commu- nications 12, 4118.

Self-propulsive active nematics 2021 Cell migration guided by long-lived spatial memory.Nature Commu- nications 12, 4118

Reference 43

Resolution: verified exact; doi, observed 2026-08-05T11:47:38.059661Z

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No event found in the named queried sources as of 2026-08-13T06:32:02.005865+00:00.

Observation 7822f5e1-fce5-47f9-bf3b-386c7ab96a2d · outbound

This paper cites 2024 Individual bacterial cells can use spatial sensing of chemical gradients to direct chemotaxis on surfaces.Nature Microbiology9, 2308–2322.

Self-propulsive active nematics 2024 Individual bacterial cells can use spatial sensing of chemical gradients to direct chemotaxis on surfaces.Nature Microbiology9, 2308–2322

Reference 44

Resolution: verified exact; doi, observed 2026-08-05T11:47:38.044700Z

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Observation 8e2ddce3-08b5-467d-8723-8669492998e8 · outbound

This paper cites 2023 On estimating the structure factor of a point process, with applications to hyperuniformity.Statistics and Comput- ing 33.

Self-propulsive active nematics 2023 On estimating the structure factor of a point process, with applications to hyperuniformity.Statistics and Comput- ing 33

Reference 45

Resolution: unresolved; no resolver link, observed 2026-08-05T11:47:37.995870Z

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