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Paper Citation Record · LEDGER

The role of gain neuromodulation in layer-5 pyramidal neurons

As of 9 August 2026, this Paper Citation Record lists 30 of 30 outbound references and 0 inbound Pith citation observations for arXiv:2507.03222.

A citation records a reference. It does not transfer a finding from one paper to another.

pith.paper-citation-record.v1
2507.03222 v2

Coverage vector

measured 30 of 30 reference resolution

Typed states for the displayed outbound observations.

Source: paper_references, paper_reference_links, observed 2026-08-06T20:20:37.743276Z

measured 30 of 30 standing notices

One-hop event checks from named stored sources.

Source: scholarly_work_events, retraction_status_cache, observed 2026-08-09T06:31:02.800959+00:00

measured 0 of 0 inbound itemization

Pith citing papers itemized under the disclosed page cap.

Source: paper_references, paper_reference_links

measured 0 of 1 external citation measurements

A source-named dated measurement, never combined with another source.

Source: cited_works

Reference resolution

30 of 30 outbound references displayed

  • verified exact18
  • verified fuzzy2
  • unresolved5
  • parse uncertain0
  • malformed identifier2
  • metadata mismatch3

External citation measurements

No source-named external measurement is stored.

Outbound references

Observation e75cb165-8fa8-4365-97e4-2159c1ab811d · outbound

This paper cites E., Zhu, J.

The role of gain neuromodulation in layer-5 pyramidal neurons E., Zhu, J

Reference 1

Resolution
malformed identifier
doi_truncated, observed 2026-08-06T20:20:40.725345Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-09T06:31:02.800959+00:00.

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Observation b6ad5d39-0cb3-4a5a-8087-69236eccf0d2 · outbound

This paper cites an unresolved cited work.

The role of gain neuromodulation in layer-5 pyramidal neurons Unresolved cited work

Reference 2

Resolution
verified exact
doi, observed 2026-08-06T20:20:40.576161Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-09T06:31:02.800959+00:00.

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Observation 67a6b1a3-65af-4d60-b34e-4de2cc34876d · outbound

This paper cites & Sejnowski, T.

The role of gain neuromodulation in layer-5 pyramidal neurons & Sejnowski, T

Reference 3

Resolution
verified exact
doi, observed 2026-08-06T20:20:40.472143Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-09T06:31:02.800959+00:00.

source=pdf_text observed=2026-08-06T20:20:35.052905Z digest=sha256:fcf3073f92504c5e9da5cfc585d95aa4514e9bdcc2ff527634cfa2b41360167e

Observation 49ce1e63-b66e-4a75-90d6-ffdb9264b2ac · outbound

This paper cites an unresolved cited work.

The role of gain neuromodulation in layer-5 pyramidal neurons Unresolved cited work

Reference 4

Resolution
verified exact
doi, observed 2026-08-06T20:20:40.331814Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-09T06:31:02.800959+00:00.

source=pdf_text observed=2026-08-06T20:20:35.189334Z digest=sha256:fbbfb194dd8cdcb1db4a79e5212d1a5a3584658c4a7547899d41e9cf1e06c255

Observation a7bef411-966e-41ad-898d-8838ace87795 · outbound

This paper cites J.et al.A burst-dependent thalamocortical substrate for perceptual awareness.PLOS Comput.

The role of gain neuromodulation in layer-5 pyramidal neurons J.et al.A burst-dependent thalamocortical substrate for perceptual awareness.PLOS Comput

Reference 5

Resolution
verified exact
doi, observed 2026-08-06T20:20:40.178533Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-09T06:31:02.800959+00:00.

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Observation 1787471d-e562-4e72-a62c-24a3fc21b0a0 · outbound

This paper cites R.et al.Neuronal connected burst cascades bridge macroscale adaptive signatures across arousal states.Nat.

The role of gain neuromodulation in layer-5 pyramidal neurons R.et al.Neuronal connected burst cascades bridge macroscale adaptive signatures across arousal states.Nat

Reference 6

Resolution
verified exact
doi, observed 2026-08-06T20:20:40.038592Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-09T06:31:02.800959+00:00.

source=pdf_text observed=2026-08-06T20:20:35.373207Z digest=sha256:558112ec00277c4873efb5ea58fb95435420253da54ece42e3162df80769aa51

Observation 38e69dd3-6c1c-4504-a58a-b810a95bd251 · outbound

This paper cites an unresolved cited work.

The role of gain neuromodulation in layer-5 pyramidal neurons Unresolved cited work

Reference 7

Resolution
unresolved
no resolver link, observed 2026-08-06T20:20:35.462843Z

Source-reported events for the cited work

Unavailable: canonical work link unavailable.

source=pdf_text observed=2026-08-06T20:20:35.462843Z digest=sha256:6729062e60e21fc4937132a6b7489e0690c73accf125c9393e8af2922a9528f2

Observation 951e1cfc-86c5-4cd9-b3e0-11a87a36eafe · outbound

This paper cites & Szabadi, E.

The role of gain neuromodulation in layer-5 pyramidal neurons & Szabadi, E

Reference 8

Resolution
verified exact
doi, observed 2026-08-06T20:20:39.845540Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-09T06:31:02.800959+00:00.

source=pdf_text observed=2026-08-06T20:20:35.568614Z digest=sha256:5809cb8eaa751de9557d2b7ece96e6c07c4e1adac012a524fcb069ae7bfd063f

Observation 5ecd881a-b157-4982-a0da-aeb7826137bc · outbound

This paper cites an unresolved cited work.

The role of gain neuromodulation in layer-5 pyramidal neurons Unresolved cited work

Reference 9

Resolution
verified exact
doi, observed 2026-08-06T20:20:39.728573Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-09T06:31:02.800959+00:00.

source=pdf_text observed=2026-08-06T20:20:35.641955Z digest=sha256:0ab0a8ce0d303ba07f6b72fe6838e4c6b204b0eb2653091896a196adbbb5778b

Observation 88c4b46a-686b-473d-b502-780feb5bec18 · outbound

This paper cites an unresolved cited work.

The role of gain neuromodulation in layer-5 pyramidal neurons Unresolved cited work

Reference 10

Resolution
verified exact
doi, observed 2026-08-06T20:20:39.595010Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-09T06:31:02.800959+00:00.

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Observation 84a5ceeb-d71b-45d5-a232-ac7359133cab · outbound

This paper cites R.et al.A thalamocortical substrate for integrated information via critical synchronous bursting.

The role of gain neuromodulation in layer-5 pyramidal neurons R.et al.A thalamocortical substrate for integrated information via critical synchronous bursting

Reference 11

Resolution
verified exact
doi, observed 2026-08-06T20:20:39.460173Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-09T06:31:02.800959+00:00.

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Observation 4736354a-81df-49b0-869b-4e2a0b5b5737 · outbound

This paper cites an unresolved cited work.

The role of gain neuromodulation in layer-5 pyramidal neurons Unresolved cited work

Reference 12

Resolution
unresolved
no resolver link, observed 2026-08-06T20:20:35.957265Z

Source-reported events for the cited work

Unavailable: canonical work link unavailable.

source=pdf_text observed=2026-08-06T20:20:35.957265Z digest=sha256:6823b5dec7c44190e35839433e4133f87edb6ebdc801d99a6b948b0512fb518a

Observation 989adac8-b572-41c5-a5db-b04d34b3cb09 · outbound

This paper cites Simple model of spiking neurons.IEEE Transactions on Neural Networks14, 1569–1572, DOI: 10.1109/TNN.2003.820440 (2003).

The role of gain neuromodulation in layer-5 pyramidal neurons Simple model of spiking neurons.IEEE Transactions on Neural Networks14, 1569–1572, DOI: 10.1109/TNN.2003.820440 (2003)

Reference 13

Resolution
metadata mismatch
raw_fallback, observed 2026-08-06T20:20:42.024770Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-09T06:31:02.800959+00:00.

source=pdf_text observed=2026-08-06T20:20:36.107678Z digest=sha256:88986f7e591b9a13bec8d934d63a7953b31e717e048f60f537c4d0249752f837

Observation 62fddfad-c1b1-426e-a535-dfb934795174 · outbound

This paper cites & Ramaswamy, S.

The role of gain neuromodulation in layer-5 pyramidal neurons & Ramaswamy, S

Reference 14

Resolution
verified exact
doi, observed 2026-08-06T20:20:39.281263Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-09T06:31:02.800959+00:00.

source=pdf_text observed=2026-08-06T20:20:36.200619Z digest=sha256:2a5ebfd40fa00810d2a0dfa4ab1e4894b04093ff2896f8c5a206f20b35b77285

Observation 5cd0c0d6-49c2-421b-ab4d-9c941ce11e31 · outbound

This paper cites an unresolved cited work.

The role of gain neuromodulation in layer-5 pyramidal neurons Unresolved cited work

Reference 15

Resolution
unresolved
no resolver link, observed 2026-08-06T20:20:36.300643Z

Source-reported events for the cited work

Unavailable: canonical work link unavailable.

source=pdf_text observed=2026-08-06T20:20:36.300643Z digest=sha256:b8270daa27344cde1c9018955d65e017668399c3c6bcd3bca39758ff970cd1c1

Observation a27ed178-49ec-477a-8890-25ae58f6c86f · outbound

This paper cites M.Dynamical Systems in Neuroscience: The Geometry of Excitability and Bursting(The MIT Press, 2006).

The role of gain neuromodulation in layer-5 pyramidal neurons M.Dynamical Systems in Neuroscience: The Geometry of Excitability and Bursting(The MIT Press, 2006)

Reference 16

Resolution
verified fuzzy
raw_fallback, observed 2026-08-06T20:20:42.881707Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-09T06:31:02.800959+00:00.

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Observation 8b720fc5-e310-4bdf-b05d-f890fe60eba6 · outbound

This paper cites & Sprekeler, H.

The role of gain neuromodulation in layer-5 pyramidal neurons & Sprekeler, H

Reference 17

Resolution
verified exact
doi, observed 2026-08-06T20:20:39.057677Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-09T06:31:02.800959+00:00.

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Observation 419b7251-e963-43ff-ac4e-7001dfff33ed · outbound

This paper cites M.et al.The stochastic nature of action potential backpropagation in apical tuft dendrites.J.

The role of gain neuromodulation in layer-5 pyramidal neurons M.et al.The stochastic nature of action potential backpropagation in apical tuft dendrites.J

Reference 18

Resolution
verified exact
raw_fallback, observed 2026-08-06T20:20:41.738995Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-09T06:31:02.800959+00:00.

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Observation 0f76b91e-c211-4f18-b47d-d45340eb224d · outbound

This paper cites Version Number: 1.

The role of gain neuromodulation in layer-5 pyramidal neurons Version Number: 1

Reference 19

Resolution
verified exact
doi, observed 2026-08-06T20:20:38.881589Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-09T06:31:02.800959+00:00.

source=pdf_text observed=2026-08-06T20:20:36.686456Z digest=sha256:5fe6969857bee268ad0dd43546dddb0aa3763699e4967b7e44a8c5c4036611a9

Observation 6830da65-7ce3-4d6b-972e-3cb5fc2d7be2 · outbound

This paper cites & Larkum, M.

The role of gain neuromodulation in layer-5 pyramidal neurons & Larkum, M

Reference 20

Resolution
verified exact
doi, observed 2026-08-06T20:20:38.655898Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-09T06:31:02.800959+00:00.

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Observation 59f6f1c2-8bc7-4a5f-8274-b07ebde76a34 · outbound

This paper cites Viewpoint: the core and matrix of thalamic organization.Neuroscience85, 331–345, DOI: 10.1016/ S0306-4522(97)00581-2 (1998).

The role of gain neuromodulation in layer-5 pyramidal neurons Viewpoint: the core and matrix of thalamic organization.Neuroscience85, 331–345, DOI: 10.1016/ S0306-4522(97)00581-2 (1998)

Reference 21

Resolution
malformed identifier
raw_fallback, observed 2026-08-06T20:20:42.568733Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-09T06:31:02.800959+00:00.

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Observation 3c736079-f09d-473d-8522-72d2f43b9e19 · outbound

This paper cites an unresolved cited work.

The role of gain neuromodulation in layer-5 pyramidal neurons Unresolved cited work

Reference 22

Resolution
verified exact
doi, observed 2026-08-06T20:20:38.508023Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-09T06:31:02.800959+00:00.

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Observation ca095f40-adc0-47a7-9824-83595a95c611 · outbound

This paper cites End-to-End Lip Synchronisation Based on Pattern Classification.

The role of gain neuromodulation in layer-5 pyramidal neurons End-to-End Lip Synchronisation Based on Pattern Classification

Reference 23

Resolution
verified exact
local_arxiv, observed 2026-08-06T20:20:41.379873Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-09T06:31:02.800959+00:00.

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Observation d5b5138a-e4f6-469b-ab14-2af0675e0468 · outbound

This paper cites Dendritic cortical microcircuits approximate the backpropagation algorithm.

The role of gain neuromodulation in layer-5 pyramidal neurons Dendritic cortical microcircuits approximate the backpropagation algorithm

Reference 24

Resolution
verified exact
local_arxiv, observed 2026-08-06T20:20:38.331081Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-09T06:31:02.800959+00:00.

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Observation 4c101e09-b2b1-436f-b872-3ec144f09a71 · outbound

This paper cites Single-phase deep learning in cortico-cortical networks.

The role of gain neuromodulation in layer-5 pyramidal neurons Single-phase deep learning in cortico-cortical networks

Reference 25

Resolution
metadata mismatch
local_arxiv, observed 2026-08-06T20:20:38.192029Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-09T06:31:02.800959+00:00.

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Observation 0885a44b-7e12-4f05-a764-d6b1bbac1232 · outbound

This paper cites Version Number: 1.

The role of gain neuromodulation in layer-5 pyramidal neurons Version Number: 1

Reference 26

Resolution
unresolved
no resolver link, observed 2026-08-06T20:20:37.407127Z

Source-reported events for the cited work

Unavailable: canonical work link unavailable.

source=pdf_text observed=2026-08-06T20:20:37.407127Z digest=sha256:f590812668fd6496dfa8afe788931f4dd2f98ac52b226084c38d48d71b300969

Observation 8bf25ff6-2588-4f22-94d3-0c4adcfccc3c · outbound

This paper cites an unresolved cited work.

The role of gain neuromodulation in layer-5 pyramidal neurons Unresolved cited work

Reference 27

Resolution
metadata mismatch
raw_fallback, observed 2026-08-06T20:20:41.133416Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-09T06:31:02.800959+00:00.

source=pdf_text observed=2026-08-06T20:20:37.496071Z digest=sha256:7bb80532cf0d2380991a716545a3b62b96c5831a33b5d0db67872c125f089062

Observation ecd638a7-b053-43d8-a10f-85e8e7db04dc · outbound

This paper cites & Ramaswamy, S.

The role of gain neuromodulation in layer-5 pyramidal neurons & Ramaswamy, S

Reference 28

Resolution
unresolved
no resolver link, observed 2026-08-06T20:20:37.587895Z

Source-reported events for the cited work

Unavailable: canonical work link unavailable.

source=pdf_text observed=2026-08-06T20:20:37.587895Z digest=sha256:6a65533fe7152941b10d0540996e0a30ac6b5d4a91eb1e8cd5420c8e2655b936

Observation 42d0941d-0b72-4e84-82e2-5f07c37a2511 · outbound

This paper cites & Trevelyan, A.

The role of gain neuromodulation in layer-5 pyramidal neurons & Trevelyan, A

Reference 29

Resolution
verified exact
doi, observed 2026-08-06T20:20:37.921359Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-09T06:31:02.800959+00:00.

source=pdf_text observed=2026-08-06T20:20:37.644098Z digest=sha256:6a1b7bfe869b7d114051b11b841f36952d8d9d7e754a153be192813f6156d3b4

Observation 348340b1-dbb6-4655-aed7-71d6f18f0001 · outbound

This paper cites slow” (tonic) and “fast.

The role of gain neuromodulation in layer-5 pyramidal neurons slow” (tonic) and “fast

Reference 30

Resolution
verified fuzzy
raw_fallback, observed 2026-08-06T20:20:42.300515Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-09T06:31:02.800959+00:00.

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Pith citing papers

No inbound Pith citation observations are available.