Typed states for the displayed outbound observations.
Source: paper_references, paper_reference_links, observed 2026-08-06T11:58:49.467988Z
Paper Citation Record · LEDGER
As of 23 August 2026, this Paper Citation Record lists 56 of 56 outbound references and 0 inbound Pith citation observations for arXiv:2507.22273.
A citation records a reference. It does not transfer a finding from one paper to another.
Typed states for the displayed outbound observations.
Source: paper_references, paper_reference_links, observed 2026-08-06T11:58:49.467988Z
One-hop event checks from named stored sources.
Source: scholarly_work_events, retraction_status_cache, observed 2026-08-22T06:32:14.747728+00:00
Pith citing papers itemized under the disclosed page cap.
Source: paper_references, paper_reference_links
A source-named dated measurement, never combined with another source.
Source: cited_works
56 of 56 outbound references displayed
External citation measurements
No source-named external measurement is stored.
Observation 78d1440a-8ea7-4144-a47f-66c712b537b8 · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles 3uQR5tyJUwsh+o6vlaMx3inE4oU=
Reference 1
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation f8381060-da92-4eb7-9a8f-b951ff4d1190 · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles The mitotic spindle: a self-made machine.Science, 294(5542):543–547, 2001
Reference 2
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation 889eb190-255f-4535-b751-5013b1ed177d · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles Force and length in the mitotic spindle.Current Biology, 19(17):R749–R761, 2009
Reference 3
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation 7b2463f2-e294-4dec-ba17-f494ee97e171 · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles Phenotypic profiling of the human genome by time-lapse microscopy reveals cell division genes.Nature, 464(7289):721–727, 2010
Reference 4
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation 447203d4-24a7-4a63-a71d-93214b46dc24 · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles Proteome analysis of the human mitotic spindle.Molecular & cellular proteomics, 4(1):35–43, 2005
Reference 5
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation b6bef3d0-227d-43cc-85e6-e57f3eb61ea9 · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles Deep proteomics of the xenopus laevis egg using an mrna-derived reference database.Current biology, 24(13):1467–1475, 2014
Reference 6
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation 86f34052-50a0-4bb5-9ae5-3abc04d71421 · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles Cambridge university press Cambridge, 1995
Reference 7
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation aa2b8de5-d8f1-46b5-b4ee-31517ca47539 · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles Asters, vortices, and rotating spirals in active gels of polar filaments.Physical review letters, 92(7):078101, 2004
Reference 8
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation 3de016cb-fdef-41bd-ae9c-8c9e3b846653 · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles The mechanics and statistics of active matter.Annu
Reference 9
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation 8d54800d-9d97-400b-8c50-2991dce382ab · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles Instabilities and pattern formation in active particle suspensions: Kinetic theory and continuum simulations.Physical Review Letters, 100(17):178103, 2008
Reference 10
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation bb30f972-7250-4483-9556-f75f1dd9c0c5 · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles Physical basis of spindle self-organization.Proceedings of the National Academy of Sciences, 111(52):18496–18500, 2014
Reference 11
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation 400fdf43-abbc-45ec-881a-1fc071b0a903 · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles The physics of the metaphase spindle.Annual review of biophysics, 47(1):655–673, 2018
Reference 12
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation ab52d1dc-e9af-44d7-a1c0-bf20d7877d90 · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles Long-range repulsion between chromosomes in mammalian oocyte spindles
Reference 13
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation 83b3f50c-fdea-49cc-a5c8-a4c00f3206e2 · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles Self-organization of kinetochore-fibers in human mitotic spindles.Elife, 11:e75458, 2022
Reference 14
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation bbbcc1c7-08c4-48d6-a2fd-e47653a328f0 · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles Insights into the micromechanical properties of the metaphase spindle.Cell, 145(7):1062–1074, 2011
Reference 15
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation 85c09369-94c3-468e-9f9e-0743d3aa3ae7 · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles Nucleoporin levels regulate cell cycle progression and phase-specific gene expression
Reference 16
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation 9e40b1ef-28df-40b2-bc16-0836753fc74a · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles Three- dimensional structure of kinetochore-fibers in human mitotic spindles.Elife, 11:e75459, 2022
Reference 17
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation 26a3c299-a2c4-4ea9-8eb7-cf37b1108453 · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles Using 3d large scale tomography to study force generation in the mitotic spindle, 2024
Reference 18
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation 4ca366de-03b6-4cf3-af81-ca5dcf9b75f0 · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles Hydrodynamics of soft active matter.Reviews of modern physics, 85(3):1143–1189, 2013
Reference 19
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation 342a2b6e-9bb2-4ac4-837f-8c5e09a1f385 · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles Mitotic spindles revisited–new insights from 3d electron microscopy
Reference 20
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation 1b7aaf8b-0328-4b37-9f11-cc8ab1548952 · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles A new view on polarization microscopy.Nature, 381(6585):811–812, 1996
Reference 21
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation a3ef5451-e851-4571-baa3-4b8b0aed36e3 · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles A reliable, noninvasive technique for spindle imaging and enucleation of mammalian oocytes.Nature biotechnology, 18(2):223–225, 2000
Reference 22
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation ecf01b92-a4f6-4db4-af8a-c464cedd2a9f · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles Birefringence of single and bundled microtubules.Biophysical journal, 74(1):645–654, 1998
Reference 23
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation 63dc6a80-7951-4dbc-8f52-5e42c6f80ecd · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles Nucleation and transport organize microtubules in metaphase spindles
Reference 24
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation 9d6b3cdc-96de-4181-8073-650f5fb50f15 · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles Autocatalytic microtubule nucleation determines the size and mass of xenopus laevis egg extract spindles.Elife, 7:e31149, 2018
Reference 25
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation d4b95928-51b8-4b66-9f03-7b276b19a6b4 · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles Branching microtubule nucleation in xenopus egg extracts mediated by augmin and tpx2.Cell, 152(4):768–777, 2013
Reference 26
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation 8dd0ffc6-7941-46a0-8b32-66a9d5562260 · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles Physical basis of large microtubule aster growth.Elife, 5:e19145, 2016
Reference 27
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation 243ea6b5-3816-4aa5-9dfb-c4336d1caf94 · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles Mitotic spindle: kinetochore fibers hold on tight to interpolar bundles
Reference 28
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation 6fdaec13-c257-4ebf-bb10-4e41689e4cb4 · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles The multifunctional spindle midzone in vertebrate cells at a glance
Reference 29
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation c4dad2d6-2c7c-445f-9dd6-e7d37b3db602 · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles Mechanisms underlying spindle assembly and robust- ness
Reference 30
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation 99470ace-f76d-4499-ab21-be977f6aaac9 · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles Microtubule sliding within the bridging fiber pushes kinetochore fibers apart to segregate chromosomes.Developmental cell, 43(1):11–23, 2017
Reference 31
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation a5441ef2-6a21-4a51-a660-33d6131800f2 · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles Overlap microtubules link sister k-fibres and balance the forces on bi-oriented kinetochores
Reference 32
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation e192cbc5-5371-4b30-807e-2b656ac8f777 · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles Kinetochore- and chromosome-driven transition of microtubules into bundles promotes spindle assembly
Reference 33
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation 9ada11da-4af2-48da-8240-3b9f16efc8f2 · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles Unresolved cited work
Reference 34
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation e5fab266-84e6-4032-b5dc-daba3da8fb83 · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles Torques within and outside the human spindle balance twist at anaphase
Reference 35
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation 8654dbc5-784a-4829-a541-a9ae29065171 · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles McDonald.Theory of simple liquids: with applications to soft matter
Reference 36
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation d4e173b7-e72e-4df1-97b3-39b7650d29f0 · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles Polewards microtubule flux in the mitotic spindle: evidence from photoactivation of fluorescence
Reference 37
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation de5464ca-ef23-4361-8092-80e7b1e47c69 · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles Self-straining of actively crosslinked microtubule networks.Nature physics, 15(12):1295–1300, 2019
Reference 38
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation 76624a6c-0028-4ef7-99b3-51467255964b · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles Measuring microtubule polarity in spindles with second-harmonic generation.Biophysical journal, 106(8):1578–1587, 2014
Reference 39
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation 72ae1bce-1e65-448a-a758-e846c5bc86ef · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles Probing microtubules polarity in mitotic spindles in situ using interferometric second harmonic generation microscopy.Scientific Reports, 7(1):6758, 2017
Reference 40
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation e2c0520f-c508-4e1f-94d4-9330db10219d · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles Digital image processing 4th edition, 2018
Reference 41
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation 1f4db186-bea7-4e0d-9b1d-00b46a678612 · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles Resampling fewer than n observations: gains, losses, and remedies for losses
Reference 42
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation 1977b986-6897-4b55-b304-089bdb4c6583 · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles A method for monitoring the collapse of plastic sections as a function of electron dose
Reference 43
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation d723d7af-68f7-45d1-b8eb-bb71c1e0586e · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles Point spread function of the polarized light field microscope.Journal of the Optical Society of America A, 39(6):1095–1103, 2022
Reference 44
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation d8cb2ed2-2354-464d-9b29-6dc0d9d043a0 · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles Assembling microtubule-based active matter
Reference 45
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation 17c3a40c-9d48-45e3-9665-c9e9c4bafbed · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles Higher- order assembly of microtubules by counterions: from hexagonal bundles to living necklaces.Proceedings of the National Academy of Sciences, 101(46):16099–16103, 2004
Reference 46
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation e0a8dae3-184f-4b7a-ad86-e4361212d3f0 · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles Structure and dynamics of motor-driven microtubule bundles.Soft matter, 20(29):5715–5723, 2024
Reference 47
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation c9046e27-c49f-4c6a-9c9b-026c01b7245f · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles Concepts and criteria of resolution.Superresolution Optical Microscopy: The Quest for Enhanced Resolution and Contrast, pages 13–32, 2020
Reference 48
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation 246f4967-7d79-4394-bc36-6144acfc10f8 · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles Elastic energies and director fields in liquid crystal droplets, i
Reference 49
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation 7126c8aa-f6e0-465c-b6b1-27caa4b03670 · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles semi-coherent
Reference 50
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation 8a4bd910-e8c0-4a48-b114-d2770fba1995 · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles 5 in the Main Text, supplemented with boundary conditions appropriate for HeLa spindles
Reference 51
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation 8662dd1a-d1f2-4c95-9913-63eaf495720a · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles S6, supplemented with appropriate boundary conditions
Reference 52
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation 72bb9c18-ed25-44c8-835d-03d2db6f8672 · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles (13) in the Main Text
Reference 53
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation b4e7d304-cb8d-45eb-aa8d-824d63bd8fb7 · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles In our computations of LC-PolScope correlation functions, however, we use an analysis box that is displaced from the central spindle (Main Text Fig
Reference 54
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation 90af5456-b8f2-423a-9a16-b201e7ebbd75 · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles This minimal theory is obtained by setting v1 = 0 in Main Text Eqns
Reference 55
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
Observation 1db72b87-02d7-4bc2-a6ba-b6c89f2a27c1 · outbound
Active Liquid Crystal Theory Explains the Collective Organization of Microtubules in Human Mitotic Spindles UNnQpAPKxWQ5hE2yAlAOxcejO4A=
Reference 56
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-22T06:32:14.747728+00:00.
No inbound Pith citation observations are available.