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Source: paper_references, paper_reference_links, observed 2026-08-16T00:21:22.661675Z
Paper Citation Record · LEDGER
As of 20 August 2026, this Paper Citation Record lists 52 of 52 outbound references and 0 inbound Pith citation observations for arXiv:2608.12090.
A citation records a reference. It does not transfer a finding from one paper to another.
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Source: paper_references, paper_reference_links, observed 2026-08-16T00:21:22.661675Z
One-hop event checks from named stored sources.
Source: scholarly_work_events, retraction_status_cache, observed 2026-08-19T06:32:44.657259+00:00
Pith citing papers itemized under the disclosed page cap.
Source: paper_references, paper_reference_links
A source-named dated measurement, never combined with another source.
Source: cited_works
52 of 52 outbound references displayed
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Observation e2b586d2-d2cc-46a4-8f06-1148551677ee · outbound
Task- and dataset-specific information in protein language models Unresolved cited work
Reference 1
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Observation a0373031-d8f1-4df2-93b1-bcc8b89fd58b · outbound
Task- and dataset-specific information in protein language models Comparative assessment of protein large language models for enzyme commission number prediction.BMC bioinfor- matics, 26(1):68, 2025
Reference 2
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Observation 3b29df39-ef54-444f-b691-d459a6c16291 · outbound
Task- and dataset-specific information in protein language models Language mod- els can identify enzymatic binding sites in pro- tein sequences.Computational and structural biotechnology journal, 23:1929–1937, 2024
Reference 3
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Observation 19f9d383-93e1-46e2-b9a3-516103b6cf6c · outbound
Task- and dataset-specific information in protein language models Iglm: Infilling language model- ing for antibody sequence design.Cell systems, 14(11):979–989, 2023
Reference 4
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Observation 66be9fb1-5ff8-4a0b-b682-d3697ca794da · outbound
Task- and dataset-specific information in protein language models Protein language models: Applications and perspectives
Reference 5
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Observation 5ec56ba0-229c-4178-87b9-ceb0f44ee156 · outbound
Task- and dataset-specific information in protein language models Understanding intermediate layers using linear classifier probes
Reference 6
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Observation 5e65019e-b04e-4107-a0de-5c93eaa6f23b · outbound
Task- and dataset-specific information in protein language models Deep residual learning for image recognition
Reference 7
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Observation 347f3399-d8dd-46ef-8981-c7ff730b424f · outbound
Task- and dataset-specific information in protein language models Imagenet: A large-scale hierarchical image database
Reference 8
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Observation a443e45f-a347-478e-80c8-954e4ca1b61b · outbound
Task- and dataset-specific information in protein language models Bert: Pre-training of deep bidirectional transformers for language un- derstanding
Reference 9
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Observation d0d4df10-5927-4a3d-8a33-525f2975ea3a · outbound
Task- and dataset-specific information in protein language models The geometry of hidden representations of large transformer models.Ad- vances in Neural Information Processing Sys- tems, 36:51234–51252, 2023
Reference 10
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Observation 5d0dcaff-3911-463d-9c45-99b05533631d · outbound
Task- and dataset-specific information in protein language models BERTology Meets Biology: Interpreting Attention in Protein Language Models
Reference 11
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Unavailable: canonical work link unavailable.
Observation 62f72406-c10e-48c5-811f-6bbf8a4b46d6 · outbound
Task- and dataset-specific information in protein language models Layer by Layer: Uncovering Hidden Representations in Language Models
Reference 12
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Unavailable: canonical work link unavailable.
Observation a2149d53-b84a-42a6-9aca-2cfb69cef741 · outbound
Task- and dataset-specific information in protein language models The underlying structures of self-attention: symmetry, directionality, and emergent dynamics in Transformer training
Reference 13
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Unavailable: canonical work link unavailable.
Observation e8f32944-4ea1-4f36-98e7-69ac7f450143 · outbound
Task- and dataset-specific information in protein language models Generative pretraining from pixels
Reference 14
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Observation bc828bb8-22de-4f9b-b73f-0d7d552c9403 · outbound
Task- and dataset-specific information in protein language models Evolutionary-scale prediction of atomic-level protein structure with a language model.Sci- ence, 379(6637):1123–1130, 2023
Reference 15
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Observation 9a6fbca4-6151-4207-a1c1-f91619c249eb · outbound
Task- and dataset-specific information in protein language models Layer probing improves kinase functional prediction with protein language models.arXiv preprint arXiv:2512.00376, 2025
Reference 16
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Observation 4927b43e-4f8c-446e-a58e-f632969c07be · outbound
Task- and dataset-specific information in protein language models Visualizing and understanding convolutional networks
Reference 17
Source-reported events for the cited work
Unavailable: canonical work link unavailable.
Observation 787edd45-84a6-490c-93e8-b5a8cf46d50b · outbound
Task- and dataset-specific information in protein language models Quantify- ing uncertainty in protein representations across models and tasks.Nature Methods, pages 1–9, 2026
Reference 18
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No event found in the named queried sources as of 2026-08-19T06:32:44.657259+00:00.
Observation affba33a-c15a-4ff1-98ed-99d92615f57f · outbound
Task- and dataset-specific information in protein language models Local fitness land- scape of the green fluorescent protein.Nature, 533(7603):397–401, 2016
Reference 19
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-19T06:32:44.657259+00:00.
Observation 9c51a8cd-0dad-4a8c-8d9e-8c4291e8266b · outbound
Task- and dataset-specific information in protein language models A comprehensive biophysical description of pairwise epistasis throughout an entire protein domain.Current biology, 24(22):2643–2651, 2014
Reference 20
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Observation 18ec8e7e-ca78-481b-a162-5cb71479aa5f · outbound
Task- and dataset-specific information in protein language models Global analysis of protein folding using mas- sively parallel design, synthesis, and testing.Sci- ence, 357(6347):168–175, 2017
Reference 21
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No event found in the named queried sources as of 2026-08-19T06:32:44.657259+00:00.
Observation 98f5cdba-1841-4feb-b522-0e6b455cdbf9 · outbound
Task- and dataset-specific information in protein language models Mega-scale experimental analysis of protein folding stability in biology and design.Nature, 620(7973):434–444, 2023
Reference 22
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-19T06:32:44.657259+00:00.
Observation 679cd716-47f9-41a1-b306-fb4896a38368 · outbound
Task- and dataset-specific information in protein language models Large language models generate functional protein se- quences across diverse families.Nature biotech- nology, 41(8):1099–1106, 2023
Reference 23
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-19T06:32:44.657259+00:00.
Observation f395fae3-9f83-4bb2-bd55-0fc44dd02ec7 · outbound
Task- and dataset-specific information in protein language models Meltome atlas—thermal proteome stability across the tree of life.Nature methods, 17(5):495–503, 2020
Reference 24
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-19T06:32:44.657259+00:00.
Observation 24f4290b-2258-4710-8b3a-8534c969ddd2 · outbound
Task- and dataset-specific information in protein language models Deepsol: a deep learning framework for sequence-based protein solubility prediction.Bioinformatics, 34(15):2605–2613, 2018
Reference 25
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-19T06:32:44.657259+00:00.
Observation cdcafc98-16db-41b2-b238-b35f141b23fa · outbound
Task- and dataset-specific information in protein language models Deeploc 2.0: multi-label subcellular localization prediction using protein language models.Nucleic acids research, 50(W1):W228–W234, 2022
Reference 26
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-19T06:32:44.657259+00:00.
Observation 095ce127-bc6a-4f5b-a75e-f67543c31313 · outbound
Task- and dataset-specific information in protein language models Unresolved cited work
Reference 27
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-19T06:32:44.657259+00:00.
Observation 7ac8219f-c06e-4e3e-b970-3fa94f85d856 · outbound
Task- and dataset-specific information in protein language models Protein embeddings and deep learning pre- dict binding residues for various ligand classes
Reference 28
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No event found in the named queried sources as of 2026-08-19T06:32:44.657259+00:00.
Observation eec03d69-071c-4580-b4bd-ee33c1573289 · outbound
Task- and dataset-specific information in protein language models Unresolved cited work
Reference 29
Source-reported events for the cited work
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Observation b84c58da-917e-4e2d-83f5-6ea38a859b11 · outbound
Task- and dataset-specific information in protein language models ESM Cambrian: Revealing the mysteries of proteins with unsupervised learn- ing.EvolutionaryScale Website, 12 2024
Reference 30
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No event found in the named queried sources as of 2026-08-19T06:32:44.657259+00:00.
Observation 79d08c83-8a84-4280-b843-c20f2a796559 · outbound
Task- and dataset-specific information in protein language models Prottrans: to- ward understanding the language of life through self-supervised learning.IEEE transactions on pattern analysis and machine intelligence, 44(10):7112–7127, 2021
Reference 31
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Observation ab0fa540-dd3a-43a3-a0a4-786644be7b19 · outbound
Task- and dataset-specific information in protein language models Bilingual language model for protein sequence and structure.NAR Genomics and Bioinformat- ics, 6(4):lqae150, 2024
Reference 32
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No event found in the named queried sources as of 2026-08-19T06:32:44.657259+00:00.
Observation 827f0015-d9cb-4368-a9f6-f3d668d5924e · outbound
Task- and dataset-specific information in protein language models Progen2: exploring the boundaries of protein language models.Cell systems, 14(11):968–978, 2023
Reference 33
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-19T06:32:44.657259+00:00.
Observation 0da90fa4-31d1-4ecb-89b6-75c94efe179f · outbound
Task- and dataset-specific information in protein language models Protgpt2 is a deep unsupervised lan- guage model for protein design.Nature commu- nications, 13(1):4348, 2022
Reference 34
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No event found in the named queried sources as of 2026-08-19T06:32:44.657259+00:00.
Observation dc7a5a5d-2b50-49c3-a959-a369965e136f · outbound
Task- and dataset-specific information in protein language models A new algorithm for data com- pression.The C Users Journal, 12(2):23–38, 1994
Reference 35
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No event found in the named queried sources as of 2026-08-19T06:32:44.657259+00:00.
Observation 94dbf839-807e-42fd-9695-17a334e5a92e · outbound
Task- and dataset-specific information in protein language models Attention is all you need.Advances in neural information processing systems, 30, 2017
Reference 36
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Unavailable: canonical work link unavailable.
Observation b82ec052-1763-4f1e-a2a1-247daaa0045c · outbound
Task- and dataset-specific information in protein language models Exploring the limits of transfer learning with a unified text- to-text transformer.Journal of machine learning research, 21(140):1–67, 2020
Reference 37
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Unavailable: canonical work link unavailable.
Observation b01ff8e6-3a95-42d8-9f72-49bd30b1f195 · outbound
Task- and dataset-specific information in protein language models Improving language understanding by generative pre-training, 2018
Reference 38
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-19T06:32:44.657259+00:00.
Observation 505a94e6-87b6-4391-b8c7-4cdf28040293 · outbound
Task- and dataset-specific information in protein language models Estimating the intrinsic dimension of datasets by a minimal neighbor- hood information.Scientific reports, 7(1):12140, 2017
Reference 39
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-19T06:32:44.657259+00:00.
Observation c7e8de6a-1124-4b18-bdbb-ee0359f5db26 · outbound
Task- and dataset-specific information in protein language models Hierarchical nucleation in deep neural networks.Advances in Neural Information Processing Systems, 33:7526–7536, 2020
Reference 40
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-19T06:32:44.657259+00:00.
Observation 57120472-2682-4746-81d2-b1f5e2ce2285 · outbound
Task- and dataset-specific information in protein language models Unresolved cited work
Reference 41
Source-reported events for the cited work
Unavailable: canonical work link unavailable.
Observation c55c9f16-10be-4c00-8cf9-25f2dd9193fd · outbound
Task- and dataset-specific information in protein language models Supervised learning of protein melting temperature: Cross-species vs
Reference 42
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-19T06:32:44.657259+00:00.
Observation 2333d5cc-af7b-426c-9024-0b885ce5865a · outbound
Task- and dataset-specific information in protein language models Data splitting to avoid information leakage with datasail.Nature Communications, 16(1):3337, 2025
Reference 43
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-19T06:32:44.657259+00:00.
Observation 038a28ee-0d4c-4795-9a68-e352c6b85eb9 · outbound
Task- and dataset-specific information in protein language models Bert rediscovers the classical nlp pipeline
Reference 44
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Unavailable: canonical work link unavailable.
Observation 7079a065-90e6-4582-9f3f-37b07189fba0 · outbound
Task- and dataset-specific information in protein language models Uniprot: the universal protein knowledgebase in 2025.Nucleic acids research, 52(D 1):D609– D617, 2024
Reference 45
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No event found in the named queried sources as of 2026-08-19T06:32:44.657259+00:00.
Observation 702b9f0f-3f7e-4e3d-88cd-14993b7b4dc0 · outbound
Task- and dataset-specific information in protein language models Mgnify: the microbiome sequence data analy- sis resource in 2023.Nucleic acids research, 51(D1):D753–D759, 2023
Reference 46
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No event found in the named queried sources as of 2026-08-19T06:32:44.657259+00:00.
Observation d8e46ead-7a25-4d08-940e-9e656c8f5589 · outbound
Task- and dataset-specific information in protein language models The genome portal of the de- partment of energy joint genome institute: 2014 updates.Nucleic acids research, 42(D1):D26– D31, 2014
Reference 47
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-19T06:32:44.657259+00:00.
Observation c4204ce7-ed48-4e50-aa84-2c4e76e6257b · outbound
Task- and dataset-specific information in protein language models Protein-level assembly increases pro- tein sequence recovery from metagenomic sam- 17 ples manyfold.Nature methods, 16(7):603–606, 2019
Reference 48
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-19T06:32:44.657259+00:00.
Observation d753ca65-2053-4d42-afab-7dde942d2669 · outbound
Task- and dataset-specific information in protein language models Cd-hit: a fast program for clustering and comparing large sets of protein or nucleotide sequences.Bioinformat- ics, 22(13):1658–1659, 2006
Reference 49
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-19T06:32:44.657259+00:00.
Observation c4231b2b-06df-411c-bea7-74a50a8568ae · outbound
Task- and dataset-specific information in protein language models Evaluating protein transfer learning with tape.Advances in neural information processing systems, 32, 2019
Reference 50
Source-reported events for the cited work
Unavailable: canonical work link unavailable.
Observation e8f0e8f7-6f26-4c0e-92fb-f1baf6f94c3c · outbound
Task- and dataset-specific information in protein language models Proteingym: Large- scale benchmarks for protein fitness prediction and design.Advances in neural information processing systems, 36:64331–64379, 2023
Reference 51
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No event found in the named queried sources as of 2026-08-19T06:32:44.657259+00:00.
Observation 6a449864-71e8-4a6e-aabb-9a2d6ab84b6b · outbound
Task- and dataset-specific information in protein language models Kalinina
Reference 52
Source-reported events for the cited work
No event found in the named queried sources as of 2026-08-19T06:32:44.657259+00:00.
No inbound Pith citation observations are available.