pith. sign in
Pith Number

pith:4NEDP3BB

pith:2011:4NEDP3BBJGR2HQZ2HLYPCKYMKX
not attested not anchored not stored refs pending

Epistasis can lead to fragmented neutral spaces and contingency in evolution

Ard A. Louis, Iain G. Johnston, Steffen Schaper

arxiv:1108.1150 v2 · 2011-08-04 · q-bio.PE

Add to your LaTeX paper
\usepackage{pith}
\pithnumber{4NEDP3BBJGR2HQZ2HLYPCKYMKX}

Prints a linked badge after your title and injects PDF metadata. Compiles on arXiv. Learn more · Embed verified badge

Record completeness

1 Bitcoin timestamp
2 Internet Archive
3 Author claim open · sign in to claim
4 Citations open
5 Replications open
Portable graph bundle live · download bundle · merged state
The bundle contains the canonical record plus signed events. A mirror can host it anywhere and recompute the same current state with the deterministic merge algorithm.
Receipt and verification
First computed 2026-05-18T02:21:42.449299Z
Builder pith-number-builder-2026-05-17-v1
Signature Pith Ed25519 (pith-v1-2026-05) · public key
Schema pith-number/v1.0

Canonical hash

e34837ec2149a3a3c33a3af0f12b0c55d15823420ff38d8b62a07c89e9673b6f

Aliases

arxiv: 1108.1150 · arxiv_version: 1108.1150v2 · doi: 10.48550/arxiv.1108.1150 · pith_short_12: 4NEDP3BBJGR2 · pith_short_16: 4NEDP3BBJGR2HQZ2 · pith_short_8: 4NEDP3BB
Agent API
Verify this Pith Number yourself
curl -sH 'Accept: application/ld+json' https://pith.science/pith/4NEDP3BBJGR2HQZ2HLYPCKYMKX \
  | jq -c '.canonical_record' \
  | python3 -c "import sys,json,hashlib; b=json.dumps(json.loads(sys.stdin.read()), sort_keys=True, separators=(',',':'), ensure_ascii=False).encode(); print(hashlib.sha256(b).hexdigest())"
# expect: e34837ec2149a3a3c33a3af0f12b0c55d15823420ff38d8b62a07c89e9673b6f
Canonical record JSON
{
  "metadata": {
    "abstract_canon_sha256": "3a16ba8e3c6927bfdf8af82362d087dfc1bae5d6f1dccabeffc4ec1594335d74",
    "cross_cats_sorted": [],
    "license": "http://arxiv.org/licenses/nonexclusive-distrib/1.0/",
    "primary_cat": "q-bio.PE",
    "submitted_at": "2011-08-04T17:50:50Z",
    "title_canon_sha256": "e05ec63b5597168d2fff672b3f4452f06b65c551a7666e5b993e6bf9cc5de9be"
  },
  "schema_version": "1.0",
  "source": {
    "id": "1108.1150",
    "kind": "arxiv",
    "version": 2
  }
}