Large-scale evolution and extinction in a hierarchically structured environment

A class of models for large-scale evolution and mass extinctions is presented. These models incorporate environmental changes on all scales, from influences on a single species to global effects. This is a step towards a unified picture of mass extinctions, which enables one to study coevolutionary effects and external abiotic influences with the same means. The generic features of such models are studied in a simple version, in which all environmental changes are generated at random and without feedback from other parts of the system.