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REVIEW 4 major objections 6 minor 67 references

Tuning the random walk of active colloids

T0 review · 4 major / 6 minor · reviewed 2026-08-14 · deepseek-v4-flash

Pith's one-line read A pulsed electric field turns a rolling colloid into a programmable random walker.

desk verdict A programmable Quincke-roller random walker with real experimental chops; the core single-particle result is solid, but the 'truly random tumble' claim needs a direct distribution test and the collective exponents need error bars. read the letter →

arxiv 1908.04119 v1 pith:VT5TVTAZ submitted 2019-08-12 cond-mat.soft physics.flu-dyn

classification cond-mat.softphysics.flu-dyn
keywords Quinckerotationrun-and-tumblemotionLévywalkactivecolloidsMaxwell-Wagnerrelaxationmeansquareddisplacementcollectivedynamicsmatter
verification ladder T0 review T1 audit T2 compute T3 formal

The pith

A machine-rendered reading of the paper's core claim, the machinery that carries it, and where it could break.

The reading

This paper reports a synthetic colloid whose trajectory can be programmed to match the random walks used by swimming bacteria. A micron-scale sphere rolls in a steady electric field through the Quincke effect; switching the field off and on again makes it stop, reorient, and start a new run. By choosing how long the field stays off relative to the particle's charge-relaxation time, the authors make consecutive runs either independent or correlated, and by drawing run durations from an exponential or power-law distribution they reproduce run-and-tumble motion and Lévy walks. Single-particle mean-squared displacements and velocity autocorrelations agree with the analytic predictions for those walks, and populations of the walkers form swarms, rotating clusters, and turbulent-like aggregates reminiscent of bacterial suspensions. The payoff is a tabletop system in which the microscopic motility rule, not just the particle density, can be tuned at will.

What carries the argument

The central mechanism is the Quincke instability combined with Maxwell-Wagner polarization relaxation. Quincke rotation is the spontaneous spinning of a polarized sphere around an axis perpendicular to the applied field; because that axis is degenerate in the plane perpendicular to the field, each re-polarization can pick a new direction. The paper uses the Maxwell-Wagner time $\tau_{\mathrm{mw}}$, the exponential time scale for induced surface charge to build up or decay, as the memory knob: off-times long compared with $\tau_{\mathrm{mw}}$ erase the previous orientation, while shorter off-times leave partial polarization that biases the next run. A programmable waveform generator turns this physics into a random-walk synthesizer by drawing $\tau_R$ from a target distribution and setting pulse durations accordingly.

What would settle it

Measure the full distribution of turn angles $\Delta\theta$ for off-times with $\tau_T/\tau_{\mathrm{mw}} \gg 1$; if the histogram is not flat on the circle, or if the next run direction correlates with the previous run direction beyond the mean cosine, then the runs are not independent and the claim that any random walk can be emulated fails.

Watch

Extended reading notes

Core claim

The central claim is that the classical Quincke roller, a dielectric sphere that spins and rolls when polarized in a DC electric field, can be converted into a random walker whose every run and tumble is specified in advance by the applied voltage waveform. When the field is on, the sphere rolls straight at a speed set by the field amplitude; when the field is off, it stops and discharges on the Maxwell-Wagner time $\tau_{\mathrm{mw}}$, and when the field returns the Quincke instability selects a new rotation axis. If the off-time $\tau_T$ is much larger than $\tau_{\mathrm{mw}}$, the new direction is stated to be fully randomized and the run and turn phases are independent; tuning $\tau_T/\tau_{\mathrm{mw}}$ near or below 2 introduces a controlled directional memory. Drawing run durations $\tau_R$ from a chosen probability distribution and encoding them as pulse widths yields run-and-tumble walks (exponential $\tau_R$) and Lévy walks (power-law $\tau_R$), with measured mean-squared displacement and velocity autocorrelation matching the analytic expressions for constant-speed walkers with finite turning time. The paper further reports that populations of these walkers reproduce collective signatures of bacterial suspensions, including anomalous number fluctuations and an energy spectrum scaling of $-8/3$.

Load-bearing premise

The load-bearing premise is that a fully depolarized colloid tumbles by picking its next direction uniformly at random; the reported evidence is only that the average cosine of the turning angle is near zero, which does not distinguish a uniform distribution from symmetric but non-uniform ones.

Editorial extensions

If this is right

  • One experiment can now generate ordinary random walks, run-and-tumble walks, and Lévy walks from the same colloid, with the effective diffusion coefficient set by field amplitude and pulse timing.
  • The run speed depends only on the field amplitude, so speed and walk statistics are independently tunable.
  • Populations of these walkers show collective phases seen in bacterial suspensions—swarms, rotating clusters, polar clusters, and disordered clusters—with number fluctuations more anomalous than equilibrium and an energy spectrum with $-8/3$ scaling.
  • Because every particle runs and stops on the same clock, the system provides a controlled experimental platform for testing theories that link single-particle motility patterns to emergent collective order.
  • The same waveform approach extends to alternating speeds and to waiting-time distributions that yield anomalous subdiffusion, and to other Quincke-powered particles such as helical propellers.

Reading between the lines

Editorial extensions of the paper, not claims the author makes directly.

  • Editorial inference: if the turn-angle distribution is confirmed to be uniform, the walker could serve as a programmable random-walk generator for testing optimal-search theories, for instance whether Lévy walks beat run-and-tumble in obstacle fields, without relying on live bacteria.
  • Editorial inference: the global field clock imposes synchronized runs and stops on all particles, a feature absent in bacterial suspensions; matching bacterial clustering statistics may therefore arise from a different mechanism than biological coordination, and comparing the two could separate clock-driven from interaction-driven ordering.
  • Editorial inference: scaling the same protocol to smaller colloids or lower speeds would introduce Brownian noise, yielding a controlled interpolation between the deterministic run-and-tumble regime and active Brownian motion.
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Editorial analysis

A structured set of objections, weighed in public.

Desk editor's note, referee report, and a circularity audit.

Referee Report

4 major / 6 minor

Summary. The paper reports an experimental realization of a colloidal 'Quincke roller' whose motion can be programmed as a run-and-tumble or Lévy walk by applying a pulsed DC electric field. Run durations are drawn from exponential or power-law distributions and encoded as pulse widths; the field-off interval sets the tumble time and, through the Maxwell-Wagner relaxation time, the degree of memory between runs. The authors compare measured mean-squared displacement and velocity autocorrelation functions with analytic expressions from Angelani (2013) and Detcheverry (2017) using independently measured or prescribed parameters and report good agreement. They further show that dense populations of these walkers display swarming, clustering, and turbulent-like velocity fluctuations with anomalous number fluctuations, and they argue that the platform can emulate essentially any random walk.

Significance. The single-particle results are a significant technical advance: they provide a table-top system in which run-time statistics, speed, and turn duration are independently tunable, with negligible Brownian noise (Péclet number ~10^6). The use of measured or prescribed parameters rather than fitted ones is a strength, and the agreement with published analytic MSD/VACF expressions is convincing as far as the presented observables go. If the missing tumble-angle distribution is supplied, the platform would justify the 'any random walk' claim and could serve as a testbed for theories of active matter. The collective-dynamics observations are suggestive and connect naturally to bacterial suspensions, but they are less tightly quantified than the single-particle data.

major comments (4)
  1. [Random reorientation, Fig. 1e] The randomization of the tumble is supported only by the persistence index α = ⟨cos Δθ⟩ ≈ 0 (Fig. 1e). For a renewal walk with independent runs, the theoretical MSD and VACF used in the paper (Eqs. D2–D5) depend on the run-vector covariance, which is governed by the first moment ⟨cos Δθ⟩; they cannot distinguish a uniform tumble-angle distribution from any other distribution with zero first cosine moment (e.g., symmetric ±π/2 turns with equal probability). Since the abstract's claim that the strategy can 'emulate any random walk' rests on the tumble being truly random, please report the full measured distribution of Δθ and a quantitative test of uniformity (e.g., a Kolmogorov–Smirnov test against the uniform distribution), or at minimum show that the first several Fourier harmonics of the turn-angle distribution are flat.
  2. [Appendix D, Eq. (D5)] Equation (D5) for the Lévy-walk VACF is printed as V²/(τ+τ_T) [t0^γ/((γ−1)t^{1−γ})], which scales as t^{γ−1} and has incorrect dimensions. Differentiating the Lévy MSD in Eq. (D3) via the relation stated in the text gives V²/(τ+τ_T) [t0^γ/(γ−1)] t^{1−γ}. Please correct Eq. (D5) and verify that the theoretical curve in Fig. 2i is computed with the corrected form.
  3. [Run-and-Tumble and Lévy walks, Fig. 2h] For the Lévy walk, the MSD is only described as 'consistent with' the t^{3−γ} scaling, and no fitted exponent or uncertainty is given. Because the superdiffusive exponent is the quantitative signature of a Lévy walk, please report the measured exponent from a power-law fit over the scaling regime with a confidence interval and state the fit range. The sentence 'particle's displacement follows the desired distribution' is not supported by any displayed displacement distribution; the run-time distribution is imposed by the signal and is not an output validation. Please either add the measured flight-length or displacement distribution or remove the statement.
  4. [Appendix E and Fig. 4] The cluster statistics and the anomalous number-fluctuation exponents a (cited as 0.89 and 0.84) and the energy-spectrum exponent −8/3 are presented without error bars or fit details, and the cluster definition depends on a threshold distance chosen in the range 1.4d–1.6d (Appendix E). Please provide a sensitivity analysis of the reported exponents over this threshold range and report the fitting procedure and uncertainties. This is needed to support the quantitative comparisons with bacterial suspensions.
minor comments (6)
  1. [Fig. 2 caption] 'V = 0.84 m/s' should presumably be '0.84 mm/s' to be consistent with the quoted run velocities.
  2. [Appendix C] The text refers to the 'R´ eclet number' immediately after defining the Péclet number; this should read 'Péclet number'.
  3. [Appendix E] 'S2(x1, x2) van be angularly averaged' is a typo for 'can be'.
  4. [Appendix E] 'according to it’s definition' should be 'its definition'.
  5. [p. 3] 'polysterene' should be 'polystyrene'.
  6. [Throughout] The manuscript uses a nonstandard accent in 'L´ evy'; the standard form is 'Lévy'.

Circularity Check

0 steps flagged · score 0.0 of 10

No significant circularity: the MSD/VACF agreement tests an externally derived random-walk model against measured or prescribed parameters.

full rationale

The derivation chain is self-contained with respect to the claims tested. The MSD and VACF formulas in Appendix D are taken from independent published theories (Angelani 2013 [32] and Detcheverry 2017 [62]) and are not rederived to match data. The comparison parameters V, tau, tau_T and tau_mw are either prescribed by the programmed signal (tau, tau_T), directly measured from the trajectory (V as slope of run length vs run time in Fig. 2e,j), or fixed by material properties (tau_mw), rather than fitted to the MSD/VACF curves. The only self-citations are to the authors' prior work for the experimental chamber and surfactant tuning of tau_mw ([26]), which is background and not load-bearing for the random-walk validation. The inference that full randomization follows from alpha approximately zero is weaker than a direct turn-angle distribution test, but that is an evidentiary gap, not a circular reduction: alpha is a consequence, not a definition, of uniform random reorientation. Therefore no circular step meets the required evidence standard.

Assumptions & free parameters 3 free parameters · 5 assumptions · 0 invented entities

The central result relies on standard Quincke electrohydrodynamics, published random-walk formulas, and the assumption of uniformly random reorientation after depolarization. The main designed control parameters (mean run time, Lévy exponent, cluster threshold) are experimenter-set; none are fitted to verify the walk statistics.

free parameters (3)
  • Mean run time tau = 0.075 s
    Chosen by the experimenter as the mean of the prescribed run-time distribution; used directly in the analytic MSD and VACF curves.
  • Lévy walk exponent gamma = 1.7
    Chosen by the experimenter for the power-law run-time distribution; not fitted to the resulting walk statistics.
  • Cluster identification threshold distance = 1.4d to 1.6d
    Selected by trial in Appendix E so that clusters are correctly identified; affects the cluster size statistics, number fluctuation exponents, and correlation lengths.
assumptions (5)
  • standard math Run-and-tumble and Lévy walk MSD and VACF formulas from refs. [32] and [62] are correct.
    These formulas underlie the quantitative comparisons in Figs. 2c, 2d, 2h, and 2i; the paper cites them but does not re-derive them.
  • domain assumption The Quincke rotation model (Eqs. A4-A6) describes the particle spinning and threshold field E_Q.
    The run phase is assumed to follow the standard Quincke theory from refs. [30], [58], and [59]; used to interpret the velocities and the mechanism of the tumble.
  • domain assumption The new rotation axis chosen after depolarization is uniformly random.
    This is the core assumption behind the tumble being random; supported only indirectly by the persistence index alpha about 0 in Fig. 1e.
  • domain assumption Brownian translational and rotational diffusion are negligible.
    The paper estimates Pe about 10^6 and uses this to ignore Brownian diffusion in the theoretical MSD and VACF models.
  • domain assumption During each run, speed V is constant and turn time tau_T is constant.
    The theoretical MSD and VACF expressions assume constant V and tau_T; the paper verifies constant run speed via Fig. 2e and 2j.

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Cite this review

Pith. "Pith review of Tuning the random walk of active colloids." pith.science (2026). https://pith.science/paper/VT5TVTAZ

@misc{pith2026190804119,
  author       = {Pith},
  title        = {Pith review of: Tuning the random walk of active colloids},
  year         = {2026},
  howpublished = {\url{https://pith.science/paper/VT5TVTAZ}},
  note         = {Machine review of arXiv:1908.04119}
}
read the original abstract

Active particles such as swimming bacteria or self-propelled colloids are known to spontaneously organize into fascinating large-scale dynamic structures. The emergence of these collective states from the motility pattern of the individual particles, typically a random walk, is yet to be probed in a well-defined synthetic system. Here, we report the experimental realization of intermittent colloidal motion that reproduces the run-and-tumble and Levy trajectories common to many swimming and swarming bacteria. Our strategy enables to tailor the sequence of repeated "runs" (nearly constant-speed straight-line translation) and "tumbles" (seemingly erratic turn) to emulate any random walk. This new paradigm for active locomotion at the microscale opens new opportunities for experimental explorations of the collective dynamics emerging in active suspensions. We find that population of these random walkers exhibit behaviors reminiscent of bacterial suspensions such as dynamic clusters and mesoscale turbulent-like flows.

Figures

Figures reproduced from arXiv: 1908.04119 by the authors.

Figure 1
Figure 1. FIG. 1 [PITH_FULL_IMAGE:figures/full_fig_p002_1.png] view at source ↗
Figure 2
Figure 2. d shows a sharp decay of the VACF in the case of run and tumble motion, in agreement with the theoreti￾cal predictions: V ACF(t) = V 2 e −t/τ¯ / (1 + τT /τ¯) . (4) For L´evy walk, VACF exhibits a tail (Fig. 2i) which agrees well with the theoretical prediction for the L´evy walk and shows a poor fit to an exponential curve which drops sharply to zero. This, plus the fact that particle’s displacement follows the desi… view at source ↗
Figure 3
Figure 3. FIG. 3 [PITH_FULL_IMAGE:figures/full_fig_p004_3.png] view at source ↗
Figures from the paper (3 more)
Figure 4
Figure 4. Figure 4: FIG. 4 [PITH_FULL_IMAGE:figures/full_fig_p005_4.png]
Figure 5
Figure 5. Figure 5: FIG. 5. Induced free charge distribution for a sphere with [PITH_FULL_IMAGE:figures/full_fig_p006_5.png]
Figure 6
Figure 6. Figure 6: FIG. 6. 10 second of the generated signal based on the expo [PITH_FULL_IMAGE:figures/full_fig_p007_6.png]

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Reference graph

Works this paper leans on

67 extracted references · 48 canonical work pages

  1. [1]

    Physics of microswimmers – single particle motion and col- lective behavior: a review

    J Elgeti, R G Winkler, and G Gompper. Physics of microswimmers – single particle motion and col- lective behavior: a review. Reports on Progress in Physics, 78(5):056601, 2015. URL http://stacks.iop. org/0034-4885/78/i=5/a=056601

  2. [2]

    Bacterial hydrodynamics

    Eric Lauga. Bacterial hydrodynamics. Annual Review of Fluid Mechanics, 48(1):105–130, 2016. doi: 10.1146/annurev-fluid-122414-034606. URL https:// doi.org/10.1146/annurev-fluid-122414-034606

  3. [3]

    Berg and Douglas A

    Howard C. Berg and Douglas A. Brown. Chemotaxis in escherichia coli analysed by three-dimensional tracking. Nature, 239(7):500–504, 1972

  4. [4]

    Brown, Mark C

    Teuta Pilizota, Mostyn T. Brown, Mark C. Leake, Richard W. Branch, Richard M. Berry, and Judith P. Armitage. A molecular brake, not a clutch, stops the rhodobacter sphaeroides flagellar motor. Proceedings of the National Academy of Sciences, 106(28):11582–11587,

  5. [5]

    A bacterial swimmer with two alternating speeds of propagation

    Matthias Theves, Johannes Taktikos, Vasily Zaburdaev, Holger Stark, and Carsten Beta. A bacterial swimmer with two alternating speeds of propagation. Biophysical Journal, 105(8):1915 – 1924, 2013. ISSN 0006-3495. doi:https://doi.org/10.1016/j.bpj.2013.08.047. URL http://www.sciencedirect.com/science/article/ pii/S0006349513010217

  6. [6]

    Bacterial flagellum as a propeller and as a rudder for efficient chemotaxis

    Li Xie, Tuba Altindal, Suddhashil Chattopadhyay, and Xiao-Lun Wu. Bacterial flagellum as a propeller and as a rudder for efficient chemotaxis. Proceedings of the National Academy of Sciences, 108(6):2246–2251, 2011. ISSN 0027-8424. doi:10.1073/pnas.1011953108

  7. [7]

    Ebbens and David Alexander Gre- gory

    Stephen J. Ebbens and David Alexander Gre- gory. Catalytic janus colloids: Controlling tra- jectories of chemical microswimmers. Accounts of Chemical Research, 51(9):1931–1939, 2018. doi:10.1021/acs.accounts.8b00243. URL https: //doi.org/10.1021/acs.accounts.8b00243. PMID: 30070110

  8. [8]

    Wyatt Shields IV, and Orlin D

    Koohee Han, C. Wyatt Shields IV, and Orlin D. Velev. Engineering of self-propelling microbots and microdevices powered by magnetic and elec- tric fields. Advanced Functional Materials, 28(25): 1705953, 2018. doi:10.1002/adfm.201705953. URL https://onlinelibrary.wiley.com/doi/abs/10.1002/ adfm.201705953

Show all 67 references
  1. [9]

    Bioinspired micro- robots

    Stefano Palagi and Peer Fischer. Bioinspired micro- robots. Nature Reviews Materials, 3(6):113–124, JUN 2018

  2. [10]

    Huang, F

    H.-W. Huang, F. E. Uslu, P. Katsamba, E. Lauga, M. S. Sakar, and B. J. Nelson. Adaptive locomotion of ar- tificial microswimmers. Science Advances, 5(1), 2019. doi:10.1126/sciadv.aau1532. URL https://advances. sciencemag.org/content/5/1/eaau1532

  3. [11]

    Paxton, Kevin C

    Walter F. Paxton, Kevin C. Kistler, Christine C. Olmeda, Ayusman Sen, Sarah K. St. Angelo, Yanyan Cao, Thomas E. Mallouk, Paul E. Lammert, and Vin- cent H. Crespi. Catalytic nanomotors:autonomous move- 10 ment of striped nanorods. Journal of the American Chemical Society, 126(...

  4. [12]

    Arsenault, Ian Man- ners, and Geoffrey A

    Sbastien Fournier-Bidoz, Andr C. Arsenault, Ian Man- ners, and Geoffrey A. Ozin. Synthetic self-propelled nanorotors. Chem. Commun., pages 441–443, 2005. doi: 10.1039/B414896G. URL http://dx.doi.org/10.1039/ B414896G

  5. [13]

    Howse, Richard A

    Jonathan R. Howse, Richard A. L. Jones, An- thony J. Ryan, Tim Gough, Reza Vafabakhsh, and Ramin Golestanian. Self-motile colloidal par- ticles: From directed propulsion to random walk. Phys. Rev. Lett., 99:048102, Jul 2007. doi: 10.1103/PhysRevLett.99.048102. URL https://link...

  6. [14]

    Active motion of a janus particle by self-thermophoresis in a defocused laser beam

    Hong-Ren Jiang, Natsuhiko Yoshinaga, and Masaki Sano. Active motion of a janus particle by self-thermophoresis in a defocused laser beam. Phys. Rev. Lett., 105:268302, Dec 2010. doi: 10.1103/PhysRevLett.105.268302. URL https:// link.aps.org/doi/10.1103/PhysRevLett.105.268302

  7. [15]

    Active brownian mo- tion tunable by light

    Ivo Buttinoni, Giovanni Volpe, Felix Kmmel, Giorgio Volpe, and Clemens Bechinger. Active brownian mo- tion tunable by light. Journal of Physics: Condensed Matter, 24(28):284129, 2012. URL http://stacks.iop. org/0953-8984/24/i=28/a=284129

  8. [16]

    Schmidt, and Gianaurelio Cuniberti

    Larysa Baraban, Robert Streubel, Denys Makarov, Luyang Han, Dmitriy Karnaushenko, Oliver G. Schmidt, and Gianaurelio Cuniberti. Fuel-free locomotion of janus motors: Magnetically induced thermophoresis. ACS Nano, 7(2):1360–1367, 2013. doi:10.1021/nn305726m. PMID: 23268780

  9. [17]

    Self-propulsion mech- anism of active janus particles in near-critical binary mixtures

    Sela Samin and Ren´ e van Roij. Self-propulsion mech- anism of active janus particles in near-critical binary mixtures. Phys. Rev. Lett., 115:188305, Oct 2015. doi: 10.1103/PhysRevLett.115.188305. URL https://link. aps.org/doi/10.1103/PhysRevLett.115.188305

  10. [18]

    Dynamics of self-propelled janus particles in viscoelastic fluids

    Juan Ruben Gomez-Solano, Alex Blokhuis, and Clemens Bechinger. Dynamics of self-propelled janus particles in viscoelastic fluids. Phys. Rev. Lett., 116:138301, Mar 2016. doi:10.1103/PhysRevLett.116.138301. URL https://link.aps.org/doi/10.1103/PhysRevLett. 116.138301

  11. [19]

    Memory-induced transition from a per- sistent random walk to circular motion for achiral mi- croswimmers

    N Narinder, Clemens Bechinger, and Juan Ruben Gomez-Solano. Memory-induced transition from a per- sistent random walk to circular motion for achiral mi- croswimmers. Phys. Rev. Lett., 121:078003, Aug 2018. doi:10.1103/PhysRevLett.121.078003. URL https:// link.aps.org/doi/10.11...

  12. [20]

    Stephen Ebbens, Richard A. L. Jones, Anthony J. Ryan, Ramin Golestanian, and Jonathan R. Howse. Self-assembled autonomous runners and tumblers. Phys. Rev. E, 82:015304, Jul 2010. doi: 10.1103/PhysRevE.82.015304

  13. [21]

    Ebbens, Gavin A

    Stephen J. Ebbens, Gavin A. Buxton, Alexander Alex- eev, Alireza Sadeghi, and Jonathan R. Howse. Syn- thetic running and tumbling: an autonomous naviga- tion strategy for catalytic nanoswimmers. Soft Matter, 8:3077–3082, 2012. doi:10.1039/C2SM07283A. URL http://dx.doi.org/10.1...

  14. [22]

    Optimal run-and-tumble–based transportation of a janus particle with active steer- ing

    Tomoyuki Mano, Jean-Baptiste Delfau, Junichiro Iwa- sawa, and Masaki Sano. Optimal run-and-tumble–based transportation of a janus particle with active steer- ing. Proceedings of the National Academy of Sciences, 114(13):E2580–E2589, 2017. ISSN 0027-8424. doi: 10.1073/pnas.1616013114

  15. [23]

    Run-and-tumble-like motion of active colloids in viscoelastic media

    Juan Ruben Gomez-Solano Celia Lozano and Clemens Bechinger. Run-and-tumble-like motion of active colloids in viscoelastic media. New J. Phys., 20:015008, 2018

  16. [24]

    Emergence of macroscopic directed motion in popula- tions of motile colloids

    Antoine Bricard, Jean-Baptiste Caussin, Nicolas Desreumaux, Olivier Dauchot, and Denis Bartolo. Emergence of macroscopic directed motion in popula- tions of motile colloids. Nature, 503:95–98, 2013

  17. [25]

    Emergent vortices in popula- tions of colloidal rollers

    Antoine Bricard, Jean-Baptiste Caussin, Debasish Das, Charles Savoie, Vijayakumar Chikkadi, Kyohei Shitara, Oleksandr Chepizhko, Fernando Peruani, David Saintil- lan, and Denis Bartolo. Emergent vortices in popula- tions of colloidal rollers. Nature Communications, 6:7470, 2015

  18. [26]

    G. E. Pradillo, H. Karani, and Petia M. Vlahovska. Surface electroconvection instability. Soft Matter, :: 10.1039/c9sm01163c, 2019

  19. [27]

    G. Quincke. Ueber rotation em im constanten elec- trischen felde. Ann. Phys. Chem., 59:417–86, 1896

  20. [28]

    Diffusion, subdiffusion, and localization of active colloids in random post lat- tices

    Alexandre Morin, David Lopes Cardozo, Vijayakumar Chikkadi, and Denis Bartolo. Diffusion, subdiffusion, and localization of active colloids in random post lat- tices. Phys. Rev. E, 96:042611, Oct 2017. doi: 10.1103/PhysRevE.96.042611. URL https://link.aps. org/doi/10.1103/PhysRe...

  21. [29]

    Sounds and hydrodynamics of polar active fluids

    Delphine Geyer, Alexandre Morin, and Denis Bar- tolo. Sounds and hydrodynamics of polar active fluids. Nature Materials, 17(9):789–793, SEP 2018. ISSN 1476-

  22. [30]

    Lemaire and L

    E. Lemaire and L. Lobry. Chaotic behavior in electro- rotation. Physica A, 314(1-4):663–671, November 2002

  23. [31]

    Howard C Berg. E. coli in Motion. Springer Science & Business Media, 2008

  24. [32]

    Angelani

    L. Angelani. Averaged run-and-tumble walks. EPL (Europhysics Letters), 102(2):20004, 2013. URL http: //stacks.iop.org/0295-5075/102/i=2/a=20004

  25. [33]

    L´ evy walks

    V Zaburdaev, S Denisov, and J Klafter. L´ evy walks. Reviews of Modern Physics, 87(2):483, 2015

  26. [34]

    L´ evyprocesses and infinitely divisible distributions

    Ken-iti Sato, Sato Ken-Iti, and A Katok. L´ evyprocesses and infinitely divisible distributions. Cambridge univer- sity press, 1999

  27. [35]

    Stamhuis

    William Thielicke and Eize J. Stamhuis. PIVlab – to- wards user-friendly, affordable and accurate digital par- ticle image velocimetry in MATLAB. Journal of Open Research Software, 2, oct 2014. doi:10.5334/jors.bl. URL https://doi.org/10.5334%2Fjors.bl

  28. [36]

    Pair aligning improved motility of quincke rollers

    Shi Qing Lu, Bing Yue Zhang, Zhi Chao Zhang, Yan Shi, and Tian Hui Zhang. Pair aligning improved motility of quincke rollers. Soft Matter, 14:5092–5097, 2018

  29. [37]

    Großmann, P

    R. Großmann, P. Romanczuk, M. B¨ ar, and L. Schimansky-Geier. Pattern formation in ac- tive particle systems due to competing alignment interactions. The European Physical Journal Special Topics, 224(7):1325–1347, Jul 2015. ISSN 1951-6401. doi:10.1140/epjst/e2015-02462-3. URL ...

  30. [38]

    Collective motion and nonequilib- rium cluster formation in colonies of gliding bacte- ria

    Fernando Peruani, J¨ orn Starruß, Vladimir Jakovlje- vic, Lotte Søgaard-Andersen, Andreas Deutsch, and Markus B¨ ar. Collective motion and nonequilib- rium cluster formation in colonies of gliding bacte- ria. Phys. Rev. Lett., 108:098102, Feb 2012. doi: 10.1103/PhysRevLett.108...

  31. [39]

    H. P. Zhang, Avraham Beer, E.-L. Florin, and Harry L. Swinney. Collective motion and density fluctuations in bacterial colonies. Proceedings of the National Academy of Sciences, 107(31):13626–13630, 2010. ISSN 0027-8424. doi:10.1073/pnas.1001651107. URL https://www.pnas. org/co...

  32. [40]

    Dynamic clustering and chemotactic collapse of self-phoretic active parti- cles

    Oliver Pohl and Holger Stark. Dynamic clustering and chemotactic collapse of self-phoretic active parti- cles. Phys. Rev. Lett., 112:238303, Jun 2014. doi: 10.1103/PhysRevLett.112.238303. URL https://link. aps.org/doi/10.1103/PhysRevLett.112.238303

  33. [41]

    Goldstein, and John O

    Christopher Dombrowski, Luis Cisneros, Sunita Chatkaew, Raymond E. Goldstein, and John O. Kessler. Self-concentration and large-scale coherence in bacterial dynamics. Phys. Rev. Lett., 93:098103, Aug 2004. doi:10.1103/PhysRevLett.93.098103. URL https://link.aps.org/doi/10.1103...

  34. [42]

    Cisneros, Ricardo Cortez, Christopher Dom- browski, Raymond E

    Luis H. Cisneros, Ricardo Cortez, Christopher Dom- browski, Raymond E. Goldstein, and John O. Kessler. Fluid dynamics of self-propelled microorganisms, from individuals to concentrated populations. Experiments in Fluids, 43(5):737–753, Nov 2007. ISSN 1432-1114. doi: 10.1007/s0...

  35. [43]

    H. P. Zhang, Avraham Beer, Rachel S. Smith, E.- L. Florin, and Harry L. Swinney. Swarming dy- namics in bacterial colonies. EPL (Europhysics Letters), 87(4):48011, aug 2009. doi:10.1209/0295- 5075/87/48011. URL https://doi.org/10.1209% 2F0295-5075%2F87%2F48011

  36. [44]

    Cisneros, John O

    Luis H. Cisneros, John O. Kessler, Sujoy Ganguly, and Raymond E. Goldstein. Dynamics of swimming bac- teria: Transition to directional order at high concen- tration. Phys. Rev. E, 83:061907, Jun 2011. doi: 10.1103/PhysRevE.83.061907. URL https://link.aps. org/doi/10.1103/PhysR...

  37. [45]

    Wensink, J¨ orn Dunkel, Sebastian Heiden- reich, Knut Drescher, Raymond E

    Henricus H. Wensink, J¨ orn Dunkel, Sebastian Heiden- reich, Knut Drescher, Raymond E. Goldstein, Hartmut L¨ owen, and Julia M. Yeomans. Meso-scale turbulence in living fluids. Proceedings of the National Academy of Sciences, 109(36):14308–14313, 2012. ISSN 0027-8424. doi:10.10...

  38. [46]

    Wensink, Markus B¨ ar, and Raymond E

    J¨ orn Dunkel, Sebastian Heidenreich, Knut Drescher, Henricus H. Wensink, Markus B¨ ar, and Raymond E. Goldstein. Fluid dynamics of bacterial turbulence. Phys. Rev. Lett., 110:228102, May 2013. doi: 10.1103/PhysRevLett.110.228102. URL https://link. aps.org/doi/10.1103/PhysRevL...

  39. [47]

    Vortex arrays and mesoscale turbulence of self-propelled particles

    Robert Großmann, Pawel Romanczuk, Markus B¨ ar, and Lutz Schimansky-Geier. Vortex arrays and mesoscale turbulence of self-propelled particles. Phys. Rev. Lett., 113:258104, Dec 2014. doi: 10.1103/PhysRevLett.113.258104. URL https:// link.aps.org/doi/10.1103/PhysRevLett.113.258104

  40. [48]

    Ardekani

    Gaojin Li and Arezoo M. Ardekani. Collective motion of microorganisms in a viscoelastic fluid. Phys. Rev. Lett., 117:118001, Sep 2016. doi: 10.1103/PhysRevLett.117.118001. URL https://link. aps.org/doi/10.1103/PhysRevLett.117.118001

  41. [49]

    Bacterial swarming: a model system for studying dynamic self- assembly

    Matthew F Copeland and Douglas B Weibel. Bacterial swarming: a model system for studying dynamic self- assembly. Soft matter, 5(6):1174–1187, 2009

  42. [50]

    A self-organized vortex array of hydrodynamically en- trained sperm cells

    Ingmar H Riedel, Karsten Kruse, and Jonathon Howard. A self-organized vortex array of hydrodynamically en- trained sperm cells. Science, 309(5732):300–303, 2005

  43. [51]

    Swimming path statistics of an active brownian particle with time-dependent self-propulsion

    S Babel, B ten Hagen, and H L¨ owen. Swimming path statistics of an active brownian particle with time-dependent self-propulsion. Journal of Statistical Mechanics: Theory and Experiment, 2014(2):P02011,

  44. [52]

    Distortion and destruction of colloidal flocks in disordered environments

    Alexandre Morin, Nicolas Desreumaux, Jean-Baptiste Caussin, and Denis Bartolo. Distortion and destruction of colloidal flocks in disordered environments. Nature Physics, 13:63?67, 2017

  45. [53]

    The topography of the environment alters the optimal search strategy for active particles

    Giorgio Volpe and Giovanni Volpe. The topography of the environment alters the optimal search strategy for active particles. Proceedings of the National Academy of Sciences, 114(43):11350–11355, 2017

  46. [54]

    Active parti- cles powered by quincke rotation in a bulk fluid

    Debasish Das and Eric Lauga. Active parti- cles powered by quincke rotation in a bulk fluid. Phys. Rev. Lett., 122:194503, May 2019. doi: 10.1103/PhysRevLett.122.194503. URL https://link. aps.org/doi/10.1103/PhysRevLett.122.194503

  47. [55]

    J. R. Melcher and G. I. Taylor. Electrohydrodynamics - a review of role of interfacial shear stress. Annu. Rev. Fluid Mech., 1:111–146, 1969

  48. [56]

    Complex collective dynam- ics of active torque-driven colloids at inter- faces

    Alexey Snezhko. Complex collective dynam- ics of active torque-driven colloids at inter- faces. Current Opinion Coloid and Interface Sci., 21(SI):65–75, FEB 2016. ISSN 1359-0294. doi: 10.1016/j.cocis.2015.11.010

  49. [57]

    Lavrentovich

    Oleg D. Lavrentovich. Active colloids in liquid crys- tals. Current Opinion in Colloid and Interface Sci., 21 (SI):97–109, FEB 2016. ISSN 1359-0294. doi: 10.1016/j.cocis.2015.11.008

  50. [58]

    I. Turcu. Electric field induced rotation of spheres. J. Phys. A: Math. Gen., 20:3301–3307, 1987

  51. [59]

    T. B. Jones. Quincke rotation of spheres. IEEE Trans. Industry Appl., 20:845–849, 1984

  52. [60]

    Active particles in complex and crowded envi- ronments

    Clemens Bechinger, Roberto Di Leonardo, Hartmut L¨ owen, Charles Reichhardt, Giorgio Volpe, and Giovanni Volpe. Active particles in complex and crowded envi- ronments. REVIEWS OF MODERN PHYSICS, 88(4): 045006, 2016

  53. [61]

    Felix Thiel, Lutz Schimansky-Geier, and Igor M. Sokolov. Anomalous diffusion in run-and-tumble mo- tion. Phys. Rev. E, 86:021117, Aug 2012. doi: 10.1103/PhysRevE.86.021117. URL https://link.aps. org/doi/10.1103/PhysRevE.86.021117

  54. [62]

    Generalized run-and-turn mo- tions: From bacteria to L´ evy walks

    Fran ¸ cois Detcheverry. Generalized run-and-turn mo- tions: From bacteria to L´ evy walks. Phys. Rev. E, 96:012415, Jul 2017. doi:10.1103/PhysRevE.96.012415. URL https://link.aps.org/doi/10.1103/PhysRevE. 96.012415

  55. [63]

    Bracewell

    R. Bracewell. The Fourier Transform and Its Applications. McGraw-Hill, New York, 1965

  56. [64]

    Torquato

    S. Torquato. Random Heterogeneous Materials: Microstructure and Macroscopic Properties. Springer Science & Business Media, Berlin, 2013

  57. [1122]

    doi:10.1038/s41563-018-0123-4

  58. [2009]

    doi:10.1073/pnas.0813164106

    ISSN 0027-8424. doi:10.1073/pnas.0813164106

  59. [2014]

    URL http://stacks.iop.org/1742-5468/2014/ i=2/a=P02011

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