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Paper Citation Record · LEDGER

ProGen: Language Modeling for Protein Generation

As of 9 August 2026, this Paper Citation Record lists 0 of 0 outbound references and 21 inbound Pith citation observations for arXiv:2004.03497.

A citation records a reference. It does not transfer a finding from one paper to another.

pith.paper-citation-record.v1
2004.03497 v1

Coverage vector

measured 0 of 0 reference resolution

Typed states for the displayed outbound observations.

Source: paper_references, paper_reference_links

measured 21 of 21 standing notices

One-hop event checks from named stored sources.

Source: scholarly_work_events, retraction_status_cache, observed 2026-08-08T06:32:00.761636+00:00

measured 21 of 21 inbound itemization

Pith citing papers itemized under the disclosed page cap.

Source: paper_references, paper_reference_links, observed 2026-08-08T18:43:34.217184Z

measured 1 of 1 external citation measurements

A source-named dated measurement, never combined with another source.

Source: arxiv_reference, observed 2026-08-05T02:28:24.338817Z

Reference resolution

0 of 0 outbound references displayed

  • verified exact0
  • verified fuzzy0
  • unresolved0
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  • malformed identifier0
  • metadata mismatch0

External citation measurements

34
arxiv_reference, observed 2026-08-05T02:28:24.338817Z

Outbound references

No outbound reference observations are available for this paper version.

Pith citing papers

Observation 7d2c7c9d-7cda-4a93-8d24-9ce1ab93aab2 · inbound

Rethinking Attention with Performers cites this paper.

Rethinking Attention with Performers ProGen: Language Modeling for Protein Generation

Reference 139

Resolution
metadata mismatch
arxiv_id, observed 2026-05-12T09:16:14.563280Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-08T06:32:00.761636+00:00.

source=arxiv_source observed=2026-05-12T09:16:14.336570Z digest=sha256:0861c6c8fe6e6a4930b620c989611c412070cc726e0bac750f8b8aa9786239f2

Observation 598e2b66-cddf-4b9d-9b7a-23ae232a0d2d · inbound

Scaling Data-Constrained Language Models cites this paper.

Scaling Data-Constrained Language Models ProGen: Language Modeling for Protein Generation

Reference 71

Resolution
verified exact
arxiv_id, observed 2026-05-18T01:35:21.269023Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-08T06:32:00.761636+00:00.

source=pdf_text observed=2026-05-18T01:35:21.150772Z digest=sha256:d3347ce82368b65ff6759af2433279f6bb63a848409c227ba35531bb91641ec8

Observation 3bbe578f-da88-4276-a860-345c045ea0cb · inbound

A Comprehensive Review of Protein Language Models cites this paper.

A Comprehensive Review of Protein Language Models ProGen: Language Modeling for Protein Generation

Reference 66

Resolution
unresolved
no resolver link, observed 2026-08-08T18:43:34.217184Z

Source-reported events for the cited work

Unavailable: canonical work link unavailable.

source=pdf_text observed=2026-08-08T18:43:34.217184Z digest=sha256:0266b465d4faf4e675b4a428c30ac8f4ac13e54074a174eed706ba9c73aa6251

Observation 10a03715-6621-488e-8a10-c2f8a8d39aa8 · inbound

Steering Protein Family Design through Profile Bayesian Flow cites this paper.

Steering Protein Family Design through Profile Bayesian Flow ProGen: Language Modeling for Protein Generation

Reference 27

Resolution
unresolved
no resolver link, observed 2026-08-08T12:02:09.003805Z

Source-reported events for the cited work

Unavailable: canonical work link unavailable.

source=arxiv_source observed=2026-08-08T12:02:09.003805Z digest=sha256:a11072dd4f173da60ea3c43d406ebbb0f2e6e03896116caee2e02c12aed527b3

Observation 56695bcc-2b62-4709-97c1-9b757a1d1016 · inbound

Towards More Accurate Full-Atom Antibody Co-Design cites this paper.

Towards More Accurate Full-Atom Antibody Co-Design ProGen: Language Modeling for Protein Generation

Reference 2022

Resolution
unresolved
no resolver link, observed 2026-08-08T12:27:36.736845Z

Source-reported events for the cited work

Unavailable: canonical work link unavailable.

source=pdf_text observed=2026-08-08T12:27:36.736845Z digest=sha256:ac2612224d883679d13e768887683fe8a73b94e2052c7100dbb3c62c45ede459

Observation 00413c1b-bfe0-4fda-8614-38ff98e57f7b · inbound

Ankh3: Multi-Task Pretraining with Sequence Denoising and Completion Enhances Protein Representations cites this paper.

Ankh3: Multi-Task Pretraining with Sequence Denoising and Completion Enhances Protein Representations ProGen: Language Modeling for Protein Generation

Reference 9856

Resolution
unresolved
no resolver link, observed 2026-08-07T14:04:25.436032Z

Source-reported events for the cited work

Unavailable: canonical work link unavailable.

source=pdf_text observed=2026-08-07T14:04:25.436032Z digest=sha256:d93dc3ff0f785531868e0344eac57c2a0ce4f8d957414d74d56119c3c722e3b4

Observation 4f9338bd-2ed0-4dc9-80f7-327d3af7de25 · inbound

Leveraging Natural Language Processing to Unravel the Mystery of Life: A Review of NLP Approaches in Genomics, Transcriptomics, and Proteomics cites this paper.

Leveraging Natural Language Processing to Unravel the Mystery of Life: A Review of NLP Approaches in Genomics, Transcriptomics, and Proteomics ProGen: Language Modeling for Protein Generation

Reference 117

Resolution
unresolved
no resolver link, observed 2026-08-07T11:31:19.347410Z

Source-reported events for the cited work

Unavailable: canonical work link unavailable.

source=pdf_text observed=2026-08-07T11:31:19.347410Z digest=sha256:b67240c5d7e86422ec3a30837f34210eea15834a6499d73797d445ca43faaf19

Observation 372a1738-024d-4ffa-9184-691f8a377642 · inbound

Protein Inverse Folding From Structure Feedback cites this paper.

Protein Inverse Folding From Structure Feedback ProGen: Language Modeling for Protein Generation

Reference 26

Resolution
unresolved
no resolver link, observed 2026-08-07T11:17:57.085748Z

Source-reported events for the cited work

Unavailable: canonical work link unavailable.

source=pdf_text observed=2026-08-07T11:17:57.085748Z digest=sha256:c1289ae07d1c1a8dea9d9f59a75044cd14f26d2c4ddb6082abe8b992a67d9675

Observation d4301f0d-e67f-4db0-bc1f-60f5b22cdf68 · inbound

AnnoDPO: Protein Functional Annotation Learning with Direct Preference Optimization cites this paper.

AnnoDPO: Protein Functional Annotation Learning with Direct Preference Optimization ProGen: Language Modeling for Protein Generation

Reference 2025

Resolution
unresolved
no resolver link, observed 2026-08-07T05:47:21.080984Z

Source-reported events for the cited work

Unavailable: canonical work link unavailable.

source=pdf_text observed=2026-08-07T05:47:21.080984Z digest=sha256:93eee655e4ce3b81afb020f9c3bdb1c13ddbd08a64970da0e73a902b2f6e8af8

Observation 730664f7-0f81-4239-8d70-4b0965f1777f · inbound

Protriever: End-to-End Differentiable Protein Homology Search for Fitness Prediction cites this paper.

Protriever: End-to-End Differentiable Protein Homology Search for Fitness Prediction ProGen: Language Modeling for Protein Generation

Reference 2

Resolution
unresolved
no resolver link, observed 2026-08-07T05:05:11.902139Z

Source-reported events for the cited work

Unavailable: canonical work link unavailable.

source=pdf_text observed=2026-08-07T05:05:11.902139Z digest=sha256:a5bb2345a4542421fb0ba0f6899021e9a82a617bb4c6c2af795d8276dda2477b

Observation 02aaff5b-fe6a-4190-b884-f0a24ce7fd96 · inbound

DisProtEdit: Exploring Disentangled Representations for Multi-Attribute Protein Editing cites this paper.

DisProtEdit: Exploring Disentangled Representations for Multi-Attribute Protein Editing ProGen: Language Modeling for Protein Generation

Reference 23

Resolution
unresolved
no resolver link, observed 2026-08-07T00:23:25.589286Z

Source-reported events for the cited work

Unavailable: canonical work link unavailable.

source=arxiv_source observed=2026-08-07T00:23:25.589286Z digest=sha256:e1f4b350eda519af1a30330ba655a35b232da59bedfc17c5f2b7ce0b39072aaf

Observation 29eccc50-ae49-4c8d-a4dd-934ced6a2686 · inbound

Steering Protein Language Models cites this paper.

Steering Protein Language Models ProGen: Language Modeling for Protein Generation

Reference 22

Resolution
unresolved
no resolver link, observed 2026-08-06T21:10:12.052808Z

Source-reported events for the cited work

Unavailable: canonical work link unavailable.

source=pdf_text observed=2026-08-06T21:10:12.052808Z digest=sha256:52710a95a7a43e3dab9fe54c8e7fdf0947ab2a2635312befde522df30eb44be5

Observation 563902ff-d992-46df-a51d-858da2d8c7ec · inbound

Generative Artificial Intelligence in Bioinformatics: A Systematic Review of Models, Applications, and Methodological Advances cites this paper.

Generative Artificial Intelligence in Bioinformatics: A Systematic Review of Models, Applications, and Methodological Advances ProGen: Language Modeling for Protein Generation

Reference 39

Resolution
unresolved
no resolver link, observed 2026-08-03T23:59:57.264722Z

Source-reported events for the cited work

Unavailable: canonical work link unavailable.

source=pdf_text observed=2026-08-03T23:59:57.264722Z digest=sha256:ec8198cc75665dd7980f57fe5067d13eb8f134e1e7beed85683b85e511d3f7d1

Observation b7cabfe8-917b-412e-8a16-46d8daf02c00 · inbound

From Words to Amino Acids: Does the Curse of Depth Persist? cites this paper.

From Words to Amino Acids: Does the Curse of Depth Persist? ProGen: Language Modeling for Protein Generation

Reference 23

Resolution
verified exact
arxiv_id, observed 2026-05-15T19:16:30.995526Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-08T06:32:00.761636+00:00.

source=pdf_text observed=2026-05-15T19:16:01.481502Z digest=sha256:bc8ee4b5b64f4817fec30bd5fc992713ff743fc00cab7fcb7534498eb4876c50

Observation 686e20fe-e14d-45e2-9243-b8206f5c3833 · inbound

Co-Generative De Novo Functional Protein Design cites this paper.

Co-Generative De Novo Functional Protein Design ProGen: Language Modeling for Protein Generation

Reference 2

Resolution
metadata mismatch
arxiv_id, observed 2026-05-11T16:41:18.845643Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-08T06:32:00.761636+00:00.

source=arxiv_source observed=2026-05-09T15:18:49.411291Z digest=sha256:a67f07c9c9c03a6f1c2cd360ece4eb2d99b6f61218e7e8b94d73fbe0ac60362b

Observation c480b554-064d-4303-942f-d920acf8cf6e · inbound

Fundamental Trade-Offs in Multi-Bit Watermarking of Stochastic Processes cites this paper.

Fundamental Trade-Offs in Multi-Bit Watermarking of Stochastic Processes ProGen: Language Modeling for Protein Generation

Reference 4

Resolution
metadata mismatch
arxiv_id, observed 2026-05-12T07:56:25.763381Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-08T06:32:00.761636+00:00.

source=pdf_text observed=2026-05-12T01:32:57.175350Z digest=sha256:915af36e2ad0479ddab3f5abbb6635719fc92e59e89e945968cd66243ebd33c6

Observation 20abf093-4dd9-4d93-bc05-d52ce09b0f73 · inbound

ProteinOPD: Towards Effective and Efficient Preference Alignment for Protein Design cites this paper.

ProteinOPD: Towards Effective and Efficient Preference Alignment for Protein Design ProGen: Language Modeling for Protein Generation

Reference 24

Resolution
metadata mismatch
arxiv_id, observed 2026-05-12T03:36:20.103659Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-08T06:32:00.761636+00:00.

source=pdf_text observed=2026-05-12T03:36:04.905210Z digest=sha256:058af40d3a2eca7389ba195c7e95b1004b273df89454bf037fb11e8c35d80aae

Observation e34b8cc5-17ec-4f2f-b37e-4261aa57f20b · inbound

AMix-2: Establishing Protein as a Native Modality in Large Language Models cites this paper.

AMix-2: Establishing Protein as a Native Modality in Large Language Models ProGen: Language Modeling for Protein Generation

Reference 55

Resolution
metadata mismatch
arxiv_id, observed 2026-06-28T20:12:37.761483Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-08T06:32:00.761636+00:00.

source=pdf_text observed=2026-06-28T20:05:06.168933Z digest=sha256:ffb9fd7d3a2154564cea78ae6e4884dea931689fc3377da2161d58ea6dbe3314

Observation feacf29e-abd6-4f7c-80b9-6bc94b4962a9 · inbound

Demystifying Multimodal Biomolecular Co-design With Intrinsic Geodesic Coupling cites this paper.

Demystifying Multimodal Biomolecular Co-design With Intrinsic Geodesic Coupling ProGen: Language Modeling for Protein Generation

Reference 156

Resolution
metadata mismatch
arxiv_id, observed 2026-06-28T12:02:06.000398Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-08T06:32:00.761636+00:00.

source=arxiv_source observed=2026-06-28T11:58:18.626259Z digest=sha256:575deb1015002ad342312065c84205f56e3ab2b64d8115560c7cac6fbfc54f77

Observation 58ed3fa7-0c8d-48ee-83e2-7d139050cfbb · inbound

Demystifying Multimodal Biomolecular Co-design With Intrinsic Geodesic Coupling cites this paper.

Demystifying Multimodal Biomolecular Co-design With Intrinsic Geodesic Coupling ProGen: Language Modeling for Protein Generation

Reference 217

Resolution
metadata mismatch
arxiv_id, observed 2026-07-02T01:26:24.915458Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-08T06:32:00.761636+00:00.

source=arxiv_source observed=2026-06-28T11:58:18.626259Z digest=sha256:2def17b10f6615fe5a37a6aac33cfb9660eb0adceba22b584fa60ec1f16a820f

Observation cde6bd07-259c-4495-96c7-a58a0c121751 · inbound

Preference-based Antibody Expression Ranking: Scaling with Large-scale Weak Supervision cites this paper.

Preference-based Antibody Expression Ranking: Scaling with Large-scale Weak Supervision ProGen: Language Modeling for Protein Generation

Reference 10

Resolution
unresolved
no resolver link, observed 2026-08-02T09:46:04.268762Z

Source-reported events for the cited work

Unavailable: canonical work link unavailable.

source=arxiv_source observed=2026-08-02T09:46:04.268762Z digest=sha256:1155ca98abdf73a80c4083dc21eec043f26c520e3c9af30571b428db95119bc1