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Paper Citation Record · LEDGER

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond

As of 7 August 2026, this Paper Citation Record lists 54 of 54 outbound references and 0 inbound Pith citation observations for arXiv:2510.02853.

A citation records a reference. It does not transfer a finding from one paper to another.

pith.paper-citation-record.v1
2510.02853 v3

Coverage vector

measured 54 of 54 reference resolution

Typed states for the displayed outbound observations.

Source: paper_references, paper_reference_links, observed 2026-08-04T12:44:52.165988Z

measured 54 of 54 standing notices

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Source: scholarly_work_events, retraction_status_cache, observed 2026-08-07T06:34:17.273281+00:00

measured 0 of 0 inbound itemization

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54 of 54 outbound references displayed

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Outbound references

Observation 8275d2f1-7d92-48d1-b06e-bfd196520304 · outbound

This paper cites From the neuron doctrine to neural networks.Nature reviews neuroscience, 16(8):487–497, 2015.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond From the neuron doctrine to neural networks.Nature reviews neuroscience, 16(8):487–497, 2015

Reference 1

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Observation 2c554a4b-8268-4e1e-b953-0566ad3010ae · outbound

This paper cites Computation through neural population dynamics.Annual review of neuroscience, 43(1):249–275, 2020.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Computation through neural population dynamics.Annual review of neuroscience, 43(1):249–275, 2020

Reference 2

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Observation ab978ea6-b2d3-4c27-8998-8baa82b5210a · outbound

This paper cites Neural tuning and representational geometry.Nature Reviews Neuroscience, 22(11):703–718, 2021.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Neural tuning and representational geometry.Nature Reviews Neuroscience, 22(11):703–718, 2021

Reference 3

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Observation 9962b709-2ca4-4260-ace3-23973f3f4095 · outbound

This paper cites Aneuralmanifoldviewofthebrain.NatureNeuroscience, pages 1–16, 2025.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Aneuralmanifoldviewofthebrain.NatureNeuroscience, pages 1–16, 2025

Reference 4

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Observation 1be1b993-3e74-480b-9a6f-047733b40d90 · outbound

This paper cites Large-scale neural recordings call for new insights to link brain and behavior.Nature neuroscience, 25(1):11–19, 2022.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Large-scale neural recordings call for new insights to link brain and behavior.Nature neuroscience, 25(1):11–19, 2022

Reference 5

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Observation 18467938-d9fa-487d-9216-08fd120a872d · outbound

This paper cites Neural population dynamics during reaching.Nature, 487(7405):51–56, 2012.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Neural population dynamics during reaching.Nature, 487(7405):51–56, 2012

Reference 6

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Observation 3dfce388-30b2-448c-aebd-0f605f25a2f0 · outbound

This paper cites Towardstheneuralpopulationdoctrine.Currentopinioninneurobiology, 55:103–111, 2019.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Towardstheneuralpopulationdoctrine.Currentopinioninneurobiology, 55:103–111, 2019

Reference 7

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Observation 8c38a751-f79c-45be-afb3-b8d49ff0f251 · outbound

This paper cites A unifying perspective on neural manifolds and circuits for cognition.Nature Reviews Neuroscience, 24(6):363–377, 2023.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond A unifying perspective on neural manifolds and circuits for cognition.Nature Reviews Neuroscience, 24(6):363–377, 2023

Reference 8

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Observation 0db61b56-b2d3-46da-93d5-dce15e9cbaea · outbound

This paper cites Spatial representation in the entorhinal cortex.Science, 305(5688):1258–1264, 2004.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Spatial representation in the entorhinal cortex.Science, 305(5688):1258–1264, 2004

Reference 9

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Observation b8e4860c-e4bd-4130-976b-b1d239045200 · outbound

This paper cites Microstructure of a spatial map in the entorhinal cortex.Nature, 436(7052):801–806, 2005.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Microstructure of a spatial map in the entorhinal cortex.Nature, 436(7052):801–806, 2005

Reference 10

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Observation fb4253ee-9891-493c-b405-b2aae8db0d2d · outbound

This paper cites Conjunctive representation of position, direction, and velocity in entorhinal cortex.Science, 312(5774):758–762, 2006.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Conjunctive representation of position, direction, and velocity in entorhinal cortex.Science, 312(5774):758–762, 2006

Reference 11

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Observation 4c9cde59-6a1b-4812-985d-cfea2b016afc · outbound

This paper cites Toroidal topology of population activity in grid cells.Nature, 602(7895):123–128, 2022.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Toroidal topology of population activity in grid cells.Nature, 602(7895):123–128, 2022

Reference 12

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Observation 1a655883-bc8c-4a15-9a35-33204ed8d253 · outbound

This paper cites Path integration and the neural basis of the’cognitive map’.Nature Reviews Neuroscience, 7(8):663–678, 2006.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Path integration and the neural basis of the’cognitive map’.Nature Reviews Neuroscience, 7(8):663–678, 2006

Reference 13

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Observation 9722a2f2-28a3-4f89-ad23-f0f55b666505 · outbound

This paper cites Accuratepathintegrationincontinuousattractornetworkmodelsofgridcells.PLoS computational biology, 5(2):e1000291, 2009.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Accuratepathintegrationincontinuousattractornetworkmodelsofgridcells.PLoS computational biology, 5(2):e1000291, 2009

Reference 14

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Observation 6638146f-53ec-44b2-9b8a-a57d78148c7a · outbound

This paper cites Impairedpathintegrationinmicewithdisruptedgridcellfiring.Natureneuroscience,21(1):81–91,2018.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Impairedpathintegrationinmicewithdisruptedgridcellfiring.Natureneuroscience,21(1):81–91,2018

Reference 15

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Observation bd4c5522-414b-446c-be1c-5b6e1b381202 · outbound

This paper cites A metric for space.Hippocampus, 18(12):1142–1156, 2008.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond A metric for space.Hippocampus, 18(12):1142–1156, 2008

Reference 16

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Observation 27e25033-3fd7-4c46-934e-5501d7ce020a · outbound

This paper cites Are grid cells used for navigation? on local metrics, subjective spaces, and black holes.Neuron, 111(12):1858–1875, 2023.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Are grid cells used for navigation? on local metrics, subjective spaces, and black holes.Neuron, 111(12):1858–1875, 2023

Reference 17

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Observation f6112475-ef4b-45c7-8e48-3033f67e563a · outbound

This paper cites The hippocampus as a predictive map.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond The hippocampus as a predictive map

Reference 18

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Observation e7bb5a99-8201-44ad-a6fb-df7da4c5a24a · outbound

This paper cites Vector-based navigation using grid-like representations in artificial agents.Nature, 557(7705):429–433, 2018.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Vector-based navigation using grid-like representations in artificial agents.Nature, 557(7705):429–433, 2018

Reference 19

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Observation 1ca22d36-e865-4611-8d06-5d81d616614a · outbound

This paper cites Emergence of grid-like representations by training recurrent neural networks to perform spatial localization.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Emergence of grid-like representations by training recurrent neural networks to perform spatial localization

Reference 20

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Observation eb5005cc-4d11-4edc-9f9f-18b8e1b68f9a · outbound

This paper cites A unified theory for the origin of grid cells through the lens of pattern formation.Advances in neural information processing systems, 32, 2019.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond A unified theory for the origin of grid cells through the lens of pattern formation.Advances in neural information processing systems, 32, 2019

Reference 21

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Observation 1a15939e-c4a2-49e9-a0ea-ad2caea692c3 · outbound

This paper cites The tolman-eichenbaum machine: unifying space and relational memory through generalization in the hippocampal formation.Cell, 183(5):1249–1263, 2020.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond The tolman-eichenbaum machine: unifying space and relational memory through generalization in the hippocampal formation.Cell, 183(5):1249–1263, 2020

Reference 22

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Observation 9866accc-e327-45bd-b7e6-47a3ec812706 · outbound

This paper cites On path integration of grid cells: group representation and isotropic scaling.Advances in Neural Information Processing Systems, 34:28623–28635, 2021.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond On path integration of grid cells: group representation and isotropic scaling.Advances in Neural Information Processing Systems, 34:28623–28635, 2021

Reference 23

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Observation 8ea7cedf-81a6-4ff9-afad-eec207ed6803 · outbound

This paper cites Conformal Isometry of Lie Group Representation in Recurrent Network of Grid Cells.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Conformal Isometry of Lie Group Representation in Recurrent Network of Grid Cells

Reference 24

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Observation 0623f463-79e3-4816-9dc4-942de130c5fb · outbound

This paper cites Actionable Neural Representations: Grid Cells from Minimal Constraints.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Actionable Neural Representations: Grid Cells from Minimal Constraints

Reference 25

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Observation 7bbdeb7c-911b-4563-9f4d-e15fcb49a7f0 · outbound

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The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Unresolved cited work

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Observation 04e4f306-a002-4a36-ad72-6b3958814799 · outbound

This paper cites Coherently remapping toroidal cells but not grid cells are responsible for path integration in virtual agents.Iscience, 26(11), 2023.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Coherently remapping toroidal cells but not grid cells are responsible for path integration in virtual agents.Iscience, 26(11), 2023

Reference 27

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Observation ae34ff93-8000-4a64-b05f-12e72a38faca · outbound

This paper cites A unified theory for the computational and mechanistic origins of grid cells.Neuron, 111(1):121–137, 2023.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond A unified theory for the computational and mechanistic origins of grid cells.Neuron, 111(1):121–137, 2023

Reference 28

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Observation ea00b5ac-9e68-4ffe-85c4-bdaa09b8d86b · outbound

This paper cites Self-supervised learningofrepresentationsforspacegeneratesmulti-modulargridcells.AdvancesinNeuralInformationProcessing Systems, 36:23140–23157, 2023.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Self-supervised learningofrepresentationsforspacegeneratesmulti-modulargridcells.AdvancesinNeuralInformationProcessing Systems, 36:23140–23157, 2023

Reference 29

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Observation c5317943-3892-4ef3-a0c2-d0d23373be26 · outbound

This paper cites Self-supervised grid cells without path integration.bioRxiv, pages 2024–05, 2024.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Self-supervised grid cells without path integration.bioRxiv, pages 2024–05, 2024

Reference 30

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Observation ef6a82a1-7e30-4f06-b21f-b97a4f9deb03 · outbound

This paper cites On conformal isometry of grid cells: Learning distance-preserving position embedding.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond On conformal isometry of grid cells: Learning distance-preserving position embedding

Reference 31

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Observation 1c734332-7f21-4baa-824f-0a46d27fcae3 · outbound

This paper cites Some informational aspects of visual perception.Psychological review, 61(3):183, 1954.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Some informational aspects of visual perception.Psychological review, 61(3):183, 1954

Reference 32

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Observation 230b0903-5bad-4744-aa44-7036181b252e · outbound

This paper cites Possible principles underlying the transformation of sensory messages.Sensory communication, 1(01):217–233, 1961.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Possible principles underlying the transformation of sensory messages.Sensory communication, 1(01):217–233, 1961

Reference 33

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Observation 56efdc15-e39b-4d78-9b51-b2eff7908b67 · outbound

This paper cites Second-order isomorphism of internal representations: Shapes of states.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Second-order isomorphism of internal representations: Shapes of states

Reference 34

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source=pdf_text observed=2026-08-04T12:44:50.766711Z digest=sha256:8f7dbd8008f42afaf1c3df02ff42cca00f88d3f51f2a9031daea9cd791e76d7c

Observation 254d2b48-6cf0-419e-bcc8-ee83b4f6facc · outbound

This paper cites Representation is representation of similarities.Behavioral and brain sciences, 21(4):449–467, 1998.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Representation is representation of similarities.Behavioral and brain sciences, 21(4):449–467, 1998

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source=pdf_text observed=2026-08-04T12:44:50.860739Z digest=sha256:91b61ce6d336a152ec201fac4d21ea5dff29fed159533806ceee0d6ea69fab33

Observation 1f566a23-614b-46af-a4c8-a49863195c7c · outbound

This paper cites No free lunch theorems for optimization.IEEE transactions on evolutionary computation, 1(1):67–82, 1997.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond No free lunch theorems for optimization.IEEE transactions on evolutionary computation, 1(1):67–82, 1997

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source=pdf_text observed=2026-08-04T12:44:50.925182Z digest=sha256:8d4719dcdf23c6ffda8181b443022bcbb01c219bedc4af3584b01ea0ee39db05

Observation db3e8fb6-8c35-4c4d-8d2f-a84b738a935e · outbound

This paper cites Representation learning: A review and new perspectives.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Representation learning: A review and new perspectives

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source=pdf_text observed=2026-08-04T12:44:51.052528Z digest=sha256:762f56cfabbd6122b248c325c599f99e8c3f494e10d5e327f3a887fdae097d09

Observation 8378a77c-029c-42ea-819d-3e776ed113b8 · outbound

This paper cites Geometric Deep Learning: Grids, Groups, Graphs, Geodesics, and Gauges.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Geometric Deep Learning: Grids, Groups, Graphs, Geodesics, and Gauges

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Observation 78de2de9-28ed-450b-bba6-09dd69393a2f · outbound

This paper cites Getting aligned on representational alignment.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Getting aligned on representational alignment

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source=pdf_text observed=2026-08-04T12:44:51.196076Z digest=sha256:c924bee6f0614f2271397dc98efd6715aa8460d5e719b4b30db49d15dd611113

Observation fe6e8577-ec1d-4f16-88b0-d1275204816e · outbound

This paper cites What grid cells convey about rat location.Journal of Neuroscience, 28(27):6858–6871, 2008.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond What grid cells convey about rat location.Journal of Neuroscience, 28(27):6858–6871, 2008

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source=pdf_text observed=2026-08-04T12:44:51.267970Z digest=sha256:7f1a0bb9ae1e3bac01da561b79ddc195d7fbf3de7fbdc4ef9ceb41bb61d87517

Observation 3926f411-1180-4f04-bd63-3f1dd0aadc2e · outbound

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The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond PrincetonUniversity Press, 2021

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source=pdf_text observed=2026-08-04T12:44:51.335783Z digest=sha256:fd561726ba8a2f87626c727538e5b9258af49786967fb29f5206d1ec8a0c1348

Observation a2e0aa64-b91c-4d88-9258-0bddc003ae40 · outbound

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The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Cambridge University Press, 2002

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source=pdf_text observed=2026-08-04T12:44:51.401538Z digest=sha256:86a134bc486aac9c2df32ab035338bf631176d1ec0c32520681436004ccbf8da

Observation 34563d37-aaf0-4a2e-b83a-73589bcdc9ac · outbound

This paper cites Grid cells generate an analog error-correcting code for singularly precise neural computation.Nature neuroscience, 14(10):1330–1337, 2011.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Grid cells generate an analog error-correcting code for singularly precise neural computation.Nature neuroscience, 14(10):1330–1337, 2011

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source=pdf_text observed=2026-08-04T12:44:51.445382Z digest=sha256:a47cfa05abd5caea4799c2cf6150149df16a9d3ae243f0c081e6db7b85918c19

Observation 6b0efabc-70d9-464d-8cc1-ccb15ae8ed29 · outbound

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The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond The elementary geometric structure of compact lie groups

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source=pdf_text observed=2026-08-04T12:44:51.467518Z digest=sha256:fcfe8699012302b717ad9b21339c1b381a0434e263006daca70476ddf6de11a1

Observation 836e3beb-80dd-45e5-a7c2-51131997489a · outbound

This paper cites Head-directioncellsinthedeeplayersofdorsalpresubiculumoffreelymovingrats.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Head-directioncellsinthedeeplayersofdorsalpresubiculumoffreelymovingrats

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source=pdf_text observed=2026-08-04T12:44:51.533710Z digest=sha256:b9f5fe789a0cd101f54d01e81740a0acfe83d1e235d91f676df2fa10d33c40ca

Observation 17c5aa5d-7ede-42e0-80be-77d739dc9a37 · outbound

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The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Locally ordered representation of 3d space in the entorhinal cortex.Nature, 596(7872):404–409, 2021

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source=pdf_text observed=2026-08-04T12:44:51.622164Z digest=sha256:db4ae1804d76c725f720610bb32c9b94b683adfca1d535f9968c82844ad6f260

Observation 0d89e736-2b5d-4515-8ffd-938f1464eee1 · outbound

This paper cites Irregular distribution of grid cell firing fields in rats exploring a 3d volumetric space.Nature neuroscience, 24(11):1567–1573, 2021.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Irregular distribution of grid cell firing fields in rats exploring a 3d volumetric space.Nature neuroscience, 24(11):1567–1573, 2021

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source=pdf_text observed=2026-08-04T12:44:51.682020Z digest=sha256:b1967982c24878cfeadfe6d1e20d559c79fe0e4caf2d4c75e6a1398534cd677b

Observation 8a2a9158-7610-4f27-86c9-4b3e153750e9 · outbound

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The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond A model of the neural basis of the rat’s sense of direction.Advances in neural information processing systems, 7, 1994

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source=pdf_text observed=2026-08-04T12:44:51.800531Z digest=sha256:02b7f7df9ac2fc3bbddfba9584f9957fb7ca4e4ae38c3716ff37992548869d02

Observation 89208220-91cf-4056-8779-67d058adf228 · outbound

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The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Representation of spatial orientation by the intrinsic dynamics of the head-direction cell ensemble: a theory.Journal of Neuroscience, 16(6):2112–2126, 1996

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source=pdf_text observed=2026-08-04T12:44:51.889252Z digest=sha256:c35805a7a2bcc9ea021a6034012f0585834ca1ae708e86595b7348cf347084ca

Observation 81eecba8-bbc2-4e18-83e6-690e41503262 · outbound

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The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Grid cell symmetry is shaped by environmental geometry.Nature, 518(7538):232–235, 2015

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source=pdf_text observed=2026-08-04T12:44:51.943457Z digest=sha256:d46b20c8b07c482e0f37ad729adc75f965c9123ce846b9acf096389d8f53234f

Observation 7d9699ea-172e-43a0-a8eb-61b3f72fb3b6 · outbound

This paper cites Local transformations of the hippocampal cognitive map.Science, 359(6380):1143–1146, 2018.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Local transformations of the hippocampal cognitive map.Science, 359(6380):1143–1146, 2018

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source=pdf_text observed=2026-08-04T12:44:52.027294Z digest=sha256:b99b29b912a407749c0c1f0def43c504a9226cca33a60df502de8636344b0a65

Observation 7bedaf61-a424-4309-b82e-05cd2a4f43b4 · outbound

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The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Shearing-induced asymmetry in entorhinal grid cells.Nature, 518(7538):207–212, 2015

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source=pdf_text observed=2026-08-04T12:44:52.077549Z digest=sha256:810f3c692b3d1a74e215c4c22e6973b4568832d7a6a9b98bfdbc7c06fb72969a

Observation 64b955ac-8ae7-4919-80d7-c233b4483c06 · outbound

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The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Information processing with population codes.Nature Reviews Neuroscience, 1(2):125–132, 2000

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source=pdf_text observed=2026-08-04T12:44:52.135359Z digest=sha256:5911d86d4b47ce6bd8ebb067b1568193a46f2ba8a9288a4bab5b8c890dea9a81

Observation 270eacf7-4bdb-45a5-9f3d-8130495e96fd · outbound

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The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Connecting multiple spatial scales to decode the population activity of grid cells.Science Advances, 1(11):e1500816, 2015

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source=pdf_text observed=2026-08-04T12:44:52.165988Z digest=sha256:72081638d9cdb676545c7bd2d2019fbcfd937c83b3e9193598f2a894e52ef3cf

Pith citing papers

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