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Paper Citation Record · LEDGER

Proteo-R1: Reasoning Foundation Models for De Novo Protein Design

As of 4 August 2026, this Paper Citation Record lists 29 of 29 outbound references and 2 inbound Pith citation observations for arXiv:2605.02937.

A citation records a reference. It does not transfer a finding from one paper to another.

pith.paper-citation-record.v1
2605.02937 v1

Coverage vector

measured 29 of 29 reference resolution

Typed states for the displayed outbound observations.

Source: paper_references, paper_reference_links, observed 2026-05-09T19:31:31.963035Z

measured 31 of 31 standing notices

One-hop event checks from named stored sources.

Source: scholarly_work_events, retraction_status_cache, observed 2026-08-04T06:34:03.388597+00:00

measured 2 of 2 inbound itemization

Pith citing papers itemized under the disclosed page cap.

Source: paper_references, paper_reference_links, observed 2026-08-01T22:38:27.212657Z

measured 0 of 1 external citation measurements

A source-named dated measurement, never combined with another source.

Source: pith, observed 2026-07-04T10:09:44.972677Z

Reference resolution

29 of 29 outbound references displayed

  • verified exact14
  • verified fuzzy10
  • unresolved3
  • parse uncertain0
  • malformed identifier0
  • metadata mismatch2

External citation measurements

No source-named external measurement is stored.

Outbound references

Observation 0c4c6d8f-7094-4af1-aa82-e80656b7a463 · outbound

This paper cites Toward de novo protein design from natural language.bioRxiv, pp.

Proteo-R1: Reasoning Foundation Models for De Novo Protein Design Toward de novo protein design from natural language.bioRxiv, pp

Reference 1

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verified fuzzy
raw_fallback, observed 2026-05-25T21:17:11.165035Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-04T06:34:03.388597+00:00.

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Observation 8fd0ebaa-7106-41ee-895a-549e10dbce2c · outbound

This paper cites Emerging Properties in Unified Multimodal Pretraining.

Proteo-R1: Reasoning Foundation Models for De Novo Protein Design Emerging Properties in Unified Multimodal Pretraining

Reference 2

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verified exact
local_arxiv, observed 2026-05-11T15:36:13.494361Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-04T06:34:03.388597+00:00.

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Observation b9607ac9-13ee-493a-b39d-b136acf34690 · outbound

This paper cites UniIF: Unified Molecule Inverse Folding.

Proteo-R1: Reasoning Foundation Models for De Novo Protein Design UniIF: Unified Molecule Inverse Folding

Reference 3

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verified exact
arxiv_id, observed 2026-05-11T15:36:13.377238Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-04T06:34:03.388597+00:00.

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Observation e9ed6278-fbca-4540-b7e5-d3e74db9754f · outbound

This paper cites Protenix-Mini: Efficient Structure Predictor via Compact Architecture, Few-Step Diffusion and Switchable pLM.

Proteo-R1: Reasoning Foundation Models for De Novo Protein Design Protenix-Mini: Efficient Structure Predictor via Compact Architecture, Few-Step Diffusion and Switchable pLM

Reference 4

Resolution
verified exact
arxiv_id, observed 2026-05-11T15:36:13.427356Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-04T06:34:03.388597+00:00.

source=pdf_text observed=2026-05-09T19:31:31.963035Z digest=sha256:bee634475b79b1f1bae0b24236e2a0826f0ec857677581e0d29f69da0fb301e0

Observation ca1cfeb3-c30b-44cb-a936-25802ff67b9d · outbound

This paper cites ProtDAT: A Unified Framework for Protein Sequence Design from Any Protein Text Description.

Proteo-R1: Reasoning Foundation Models for De Novo Protein Design ProtDAT: A Unified Framework for Protein Sequence Design from Any Protein Text Description

Reference 5

Resolution
verified exact
arxiv_id, observed 2026-05-11T15:36:13.212265Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-04T06:34:03.388597+00:00.

source=pdf_text observed=2026-05-09T19:31:31.963035Z digest=sha256:f83399d39c2870394c971cc5279fb3720c4a883e31db09e84997edc641db8e46

Observation a5ec3bea-7f9e-4fa3-898f-9f1384accbb7 · outbound

This paper cites ProLLM: Protein Chain-of-Thoughts Enhanced LLM for Protein-Protein Interaction Prediction.

Proteo-R1: Reasoning Foundation Models for De Novo Protein Design ProLLM: Protein Chain-of-Thoughts Enhanced LLM for Protein-Protein Interaction Prediction

Reference 6

Resolution
verified exact
arxiv_id, observed 2026-05-11T15:36:13.920211Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-04T06:34:03.388597+00:00.

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Observation 7f4840b3-c8a9-49df-931d-04c59854b1ff · outbound

This paper cites UniMoMo: Unified Generative Modeling of 3D Molecules for De Novo Binder Design.

Proteo-R1: Reasoning Foundation Models for De Novo Protein Design UniMoMo: Unified Generative Modeling of 3D Molecules for De Novo Binder Design

Reference 7

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arxiv_id, observed 2026-05-11T15:41:04.878356Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-04T06:34:03.388597+00:00.

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Observation c3825c16-be69-44ac-aa0b-5a95b08bca3a · outbound

This paper cites Xiaomi mimo-vl-miloco technical report.

Proteo-R1: Reasoning Foundation Models for De Novo Protein Design Xiaomi mimo-vl-miloco technical report

Reference 8

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verified exact
arxiv_id, observed 2026-05-11T15:36:13.685332Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-04T06:34:03.388597+00:00.

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Observation 733539f3-d13e-4cf1-85e7-792c16d10660 · outbound

This paper cites an unresolved cited work.

Proteo-R1: Reasoning Foundation Models for De Novo Protein Design Unresolved cited work

Reference 9

Resolution
unresolved
raw_fallback, observed 2026-05-25T21:17:11.168353Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-04T06:34:03.388597+00:00.

source=pdf_text observed=2026-05-09T19:31:31.963035Z digest=sha256:3dc14915eb30bafd0be51bad707f49455e139d727e5e44fd453875b522d27392

Observation 3f2d514c-c53c-49ca-96d0-65bee06bd578 · outbound

This paper cites Decoupled Weight Decay Regularization.

Proteo-R1: Reasoning Foundation Models for De Novo Protein Design Decoupled Weight Decay Regularization

Reference 10

Resolution
verified exact
local_arxiv, observed 2026-05-11T15:36:13.844538Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-04T06:34:03.388597+00:00.

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Observation f0cac398-648c-4f0a-aea3-d2bb63b0c55a · outbound

This paper cites Flexible and controllable protein design by prefix-tuning large-scale protein language models.bioRxiv, pp.

Proteo-R1: Reasoning Foundation Models for De Novo Protein Design Flexible and controllable protein design by prefix-tuning large-scale protein language models.bioRxiv, pp

Reference 11

Resolution
verified fuzzy
raw_fallback, observed 2026-05-25T21:17:11.173337Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-04T06:34:03.388597+00:00.

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Observation 74acc915-3ece-4a8c-87ce-647c38f8aa4b · outbound

This paper cites ProtTeX: Structure-In-Context Reasoning and Editing of Proteins with Large Language Models.

Proteo-R1: Reasoning Foundation Models for De Novo Protein Design ProtTeX: Structure-In-Context Reasoning and Editing of Proteins with Large Language Models

Reference 12

Resolution
verified exact
arxiv_id, observed 2026-05-11T15:36:13.965296Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-04T06:34:03.388597+00:00.

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Observation 9e71a72b-185b-42dc-a2a0-3dcae1c87d6f · outbound

This paper cites Bindcraft: one-shot design of functional protein binders.bioRxiv, pp.

Proteo-R1: Reasoning Foundation Models for De Novo Protein Design Bindcraft: one-shot design of functional protein binders.bioRxiv, pp

Reference 13

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verified fuzzy
raw_fallback, observed 2026-05-25T21:17:11.177702Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-04T06:34:03.388597+00:00.

source=pdf_text observed=2026-05-09T19:31:31.963035Z digest=sha256:dbfd46b17bcc833cbbef279094cf60665c7baaf2354f1ae9073426602e592319

Observation ff246f71-4189-4a46-9355-260363a21b3e · outbound

This paper cites P., Matusovsky, O., Parsa, M.

Proteo-R1: Reasoning Foundation Models for De Novo Protein Design P., Matusovsky, O., Parsa, M

Reference 14

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verified fuzzy
raw_fallback, observed 2026-05-25T21:17:11.181809Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-04T06:34:03.388597+00:00.

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Observation 1ab63982-2493-4755-94f3-5aa044e4a08a · outbound

This paper cites Score-Based Generative Modeling through Stochastic Differential Equations.

Proteo-R1: Reasoning Foundation Models for De Novo Protein Design Score-Based Generative Modeling through Stochastic Differential Equations

Reference 15

Resolution
metadata mismatch
local_arxiv, observed 2026-05-11T15:36:13.074451Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-04T06:34:03.388597+00:00.

source=pdf_text observed=2026-05-09T19:31:31.963035Z digest=sha256:f8316c7e44720b13d6a6feb829434975c78318adad9885c4e6c85de77e3fb55a

Observation 58c7bebd-c259-4397-b70c-3780ab18b08e · outbound

This paper cites Instructpro: Natural language guided ligand-binding protein design.

Proteo-R1: Reasoning Foundation Models for De Novo Protein Design Instructpro: Natural language guided ligand-binding protein design

Reference 16

Resolution
verified exact
arxiv_id, observed 2026-05-11T15:36:13.196148Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-04T06:34:03.388597+00:00.

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Observation acadf18f-68df-498d-bc33-fb84baf98177 · outbound

This paper cites Boltzgen: Toward universal binder design.bioRxiv, pp.

Proteo-R1: Reasoning Foundation Models for De Novo Protein Design Boltzgen: Toward universal binder design.bioRxiv, pp

Reference 17

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verified fuzzy
raw_fallback, observed 2026-05-25T21:17:11.161219Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-04T06:34:03.388597+00:00.

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Observation a6bcb0b5-2d3a-4801-aa9f-66113e5c0874 · outbound

This paper cites Pxdesign: Fast, modular, and accurate de novo design of protein binders.bioRxiv, pp.

Proteo-R1: Reasoning Foundation Models for De Novo Protein Design Pxdesign: Fast, modular, and accurate de novo design of protein binders.bioRxiv, pp

Reference 18

Resolution
verified fuzzy
raw_fallback, observed 2026-05-25T21:17:11.186257Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-04T06:34:03.388597+00:00.

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Observation 43d08761-6f3f-434f-84dd-7c211e251d6d · outbound

This paper cites Diffusion probabilistic modeling of protein backbones in 3D for the motif-scaffolding problem.

Proteo-R1: Reasoning Foundation Models for De Novo Protein Design Diffusion probabilistic modeling of protein backbones in 3D for the motif-scaffolding problem

Reference 19

Resolution
metadata mismatch
arxiv_id, observed 2026-05-11T15:36:13.545271Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-04T06:34:03.388597+00:00.

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Observation c887db91-64e2-4aa4-ae6e-a71069de84d9 · outbound

This paper cites A generative foundation model for antibody design.bioRxiv, pp.

Proteo-R1: Reasoning Foundation Models for De Novo Protein Design A generative foundation model for antibody design.bioRxiv, pp

Reference 20

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verified fuzzy
raw_fallback, observed 2026-05-25T21:17:11.148627Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-04T06:34:03.388597+00:00.

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Observation 790112d7-b963-4296-8a1e-622b4a9e4837 · outbound

This paper cites Qwen3 Technical Report.

Proteo-R1: Reasoning Foundation Models for De Novo Protein Design Qwen3 Technical Report

Reference 21

Resolution
verified exact
local_arxiv, observed 2026-05-11T15:36:13.264583Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-04T06:34:03.388597+00:00.

source=pdf_text observed=2026-05-09T19:31:31.963035Z digest=sha256:7d5d53e0fb8da425e1358fd0eaecabbac05834aee371bc752727be42c253c0cb

Observation e144ed2c-1e18-4554-a833-7667c1efe55f · outbound

This paper cites ProteinBench: A Holistic Evaluation of Protein Foundation Models.

Proteo-R1: Reasoning Foundation Models for De Novo Protein Design ProteinBench: A Holistic Evaluation of Protein Foundation Models

Reference 22

Resolution
verified exact
arxiv_id, observed 2026-05-11T15:36:13.326182Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-04T06:34:03.388597+00:00.

source=pdf_text observed=2026-05-09T19:31:31.963035Z digest=sha256:a620fedcbff705959de24913a304715ddc746695e11fc9917c75216ec8fcdd8b

Observation 43ed63cc-2bbc-44e2-917d-7ca83365fd78 · outbound

This paper cites High-affinity protein binder design via flow matching and in silico maturation.bioRxiv, pp.

Proteo-R1: Reasoning Foundation Models for De Novo Protein Design High-affinity protein binder design via flow matching and in silico maturation.bioRxiv, pp

Reference 23

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verified fuzzy
raw_fallback, observed 2026-05-25T21:17:11.144496Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-04T06:34:03.388597+00:00.

source=pdf_text observed=2026-05-09T19:31:31.963035Z digest=sha256:adf3a8e0e52180b2a48763612c92f7f68f6b7f0a1c7625644a2e01b2abaad911

Observation 63bb2855-8794-4b41-8f27-8fe2d83c0db4 · outbound

This paper cites De novo design of high-affinity protein binders with AlphaProteo.

Proteo-R1: Reasoning Foundation Models for De Novo Protein Design De novo design of high-affinity protein binders with AlphaProteo

Reference 24

Resolution
verified exact
arxiv_id, observed 2026-05-11T15:36:13.207553Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-04T06:34:03.388597+00:00.

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Observation 77c8d7d5-a551-4dfe-9b78-09df33141d65 · outbound

This paper cites CMADiff: Cross-Modal Aligned Diffusion for Controllable Protein Generation.

Proteo-R1: Reasoning Foundation Models for De Novo Protein Design CMADiff: Cross-Modal Aligned Diffusion for Controllable Protein Generation

Reference 25

Resolution
verified exact
arxiv_id, observed 2026-05-27T02:05:02.644539Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-04T06:34:03.388597+00:00.

source=pdf_text observed=2026-05-09T19:31:31.963035Z digest=sha256:adb15c1213d7762bd2bce00193ea9067a19626d59e7225c9caadbeaa0fa5002f

Observation ca42cec4-6e19-4fbd-8c94-ce4b1067cc48 · outbound

This paper cites Related Work Protein Binder and Antibody Design.Protein–protein interactions (PPIs) underlie most cellular processes and represent a major class of therapeutic targets.

Proteo-R1: Reasoning Foundation Models for De Novo Protein Design Related Work Protein Binder and Antibody Design.Protein–protein interactions (PPIs) underlie most cellular processes and represent a major class of therapeutic targets

Reference 26

Resolution
verified fuzzy
raw_fallback, observed 2026-05-25T21:17:11.156766Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-04T06:34:03.388597+00:00.

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Observation 8be58cc0-880e-47ae-8d86-59de461c7b0e · outbound

This paper cites an unresolved cited work.

Proteo-R1: Reasoning Foundation Models for De Novo Protein Design Unresolved cited work

Reference 27

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unresolved
raw_fallback, observed 2026-05-25T21:17:11.135745Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-04T06:34:03.388597+00:00.

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Observation f239ef1b-99f9-4ace-9de8-2105d27eea9a · outbound

This paper cites an unresolved cited work.

Proteo-R1: Reasoning Foundation Models for De Novo Protein Design Unresolved cited work

Reference 28

Resolution
unresolved
raw_fallback, observed 2026-05-25T21:17:11.139506Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-04T06:34:03.388597+00:00.

source=pdf_text observed=2026-05-09T19:31:31.963035Z digest=sha256:9dbbe111d04eff444b85c1858ec2808be9f794c9c51b042f98eda3c84de38f67

Observation 50d33903-6731-43d4-a317-405e0de1ab72 · outbound

This paper cites Recently, DL maturation methods similarly condition generation on predefined anchors or interaction patterns (Yu et al., 2026).

Proteo-R1: Reasoning Foundation Models for De Novo Protein Design Recently, DL maturation methods similarly condition generation on predefined anchors or interaction patterns (Yu et al., 2026)

Reference 29

Resolution
verified fuzzy
raw_fallback, observed 2026-05-25T21:17:11.152164Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-04T06:34:03.388597+00:00.

source=pdf_text observed=2026-05-09T19:31:31.963035Z digest=sha256:3dbe01846d165609fb32623b6fe18dd492273b6ca7feb7c6f9e46eb52d2927a5

Pith citing papers

Observation 055a7c6c-d60d-4ae0-a80a-8428d1b53930 · inbound

Deciphering Fingerprints of 3D Molecular Surfaces for Accurate Epitope Prediction cites this paper.

Deciphering Fingerprints of 3D Molecular Surfaces for Accurate Epitope Prediction Proteo-R1: Reasoning Foundation Models for De Novo Protein Design

Reference 86

Resolution
verified exact
local_arxiv, observed 2026-07-04T10:09:44.974100Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-04T06:34:03.388597+00:00.

source=pdf_text observed=2026-06-26T09:06:48.951623Z digest=sha256:ae356a38da0c8c5e6ccae2b604eef331d1ec6a84fbb3c637fcf80eb0023815dd

Observation 95a258f1-e515-437a-8bd5-d5856505ff91 · inbound

S1-Omni: A Unified Multimodal Reasoning Model for Scientific Understanding, Prediction, and Generation cites this paper.

S1-Omni: A Unified Multimodal Reasoning Model for Scientific Understanding, Prediction, and Generation Proteo-R1: Reasoning Foundation Models for De Novo Protein Design

Reference 31

Resolution
unresolved
no resolver link, observed 2026-08-01T22:38:27.212657Z

Source-reported events for the cited work

Unavailable: canonical work link unavailable.

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