Pith. sign in

REVIEW 3 major objections 5 minor 83 references

Activity-enhanced shear thinning of flexible linear polar polymers

T0 review · 3 major / 5 minor · reviewed 2026-08-07 · deepseek-v4-flash

Challenge this review Re-run · record.json Download PDF Read on arXiv ↗ Review JSON

Pith's one-line read Tangentially propelled polar polymers under shear flow shear-thin with viscosity exponent 4/3 and tumble with exponent 1/3 until shear dominates.

desk verdict Genuinely new active-polar-polymer shear exponents, and the simulations look mostly careful, but the viscosity route has an origin-dependence problem that needs a coordinate-invariance check before the 4/3 exponent is trusted. read the letter →

arxiv 2505.17539 v1 pith:YEDJEYRW submitted 2025-05-23 cond-mat.soft

Arindam Panda , Roland G. Winkler , Sunil P. Singh This is my paper · ORCID
classification cond-mat.soft
keywords activepolymersshearthinningtumblingdynamicspolaractivityWeissenbergnumberPécletintrinsicviscosityexcludedvolume
verification ladder T0 review T1 audit T2 compute T3 formal

The pith

A machine-rendered reading of the paper's core claim, the machinery that carries it, and where it could break.

The reading

This paper asks what happens when a flexible polymer is actively propelled along its own contour while a linear shear flow is applied. Combining coarse-grained simulations with an analytical model, it establishes a new activity-dominated rheological regime: the polymer's intrinsic viscosity falls as $\eta_p/\eta_p^0 \sim Wi_{Pe}^{-4/3}$ with shear rate, far steeper than the $Wi_{Pe}^{-1/2}$ of a passive chain, while its tumbling frequency grows only as $\nu_t \tau_r \sim Wi_{Pe}^{1/3}$ rather than $Wi_{Pe}^{2/3}$. The mechanism is that polar activity amplifies shear-induced compression in the gradient direction, so the polymer presents a smaller cross-section to the flow and overturns more slowly. At sufficiently high shear rates, ordinary flow dominates and the passive exponents return. If the claim is right, tangential polar activity is a generic route to much stronger shear thinning in dilute polymer solutions.

What carries the argument

The load-bearing machinery is a coarse-grained overdamped Langevin dynamics in which each monomer experiences an active force $f_a(\hat{t}_{i+1}+\hat{t}_i)/2$ along the bond directions together with a linear shear flow and excluded-volume forces. The companion analytical model solves a discretized Gaussian active polar polymer by expanding in the eigenfunctions of a non-symmetric matrix; its eigenvalues $\xi_n$ encode the activity-dependent mode relaxation, and both the gyration tensor and the virial stress (hence $\eta_p$) are written as sums over these modes. The paper's central scaling argument is geometric: during a tumbling event a monomer is convected at velocity $v_x \approx \dot{\gamma} y_t$ with $y_t \approx \sqrt{\langle G_{yy}\rangle}$, so $\tau_t \sim (\dot{\gamma}\sqrt{\langle G_{yy}\rangle})^{-1}$; inserting $\langle G_{yy}\rangle \sim Wi_{Pe}^{-4/3}$ gives $\nu_t \tau_r \sim Wi_{Pe}^{1/3}$. Here $Wi_{Pe} = \dot{\gamma}\tau_r(Pe)$ is the Weissenberg number built on the activity-dependent relaxation time and $Pe = f_a l_0/(k_B T)$ measures the active force.

What would settle it

Measure the intrinsic viscosity and tumbling frequency of a dilute suspension of tangentially propelled flexible polar filaments (or simulate the same model with full hydrodynamic interactions) over $10 < Wi_{Pe} < 1000$ at $Pe > 5$; finding viscosity slopes other than $-4/3$ or tumbling slopes other than $1/3$ in the window where passive chains show $-1/2$ and $2/3$ would settle the claim negatively. A sharper internal check is the paper's prediction that $\langle G_{yy}\rangle$ and $\eta_p$ track each other within 25–35% across four decades of $Wi_{Pe}$; a measurement where those two curves diverge would falsify the proposed mechanism.

Watch

Extended reading notes

Core claim

The central discovery is a qualitative change in the shear-rate scaling of dilute tangentially driven active polar polymers. In the activity-dominated intermediate window, the normalized intrinsic viscosity obeys $\eta_p/\eta_p^0 \sim Wi_{Pe}^{-4/3}$, the gradient-direction radius of gyration shrinks as $\langle G_{yy}\rangle/\langle G_{yy}^0\rangle \sim Wi_{Pe}^{-4/3}$, the flow alignment satisfies $\tan(2\chi) \sim Wi_{Pe}^{-1}$, and the tumbling frequency obeys $\nu_t \tau_r \sim Wi_{Pe}^{1/3}$; all quantities then cross over to the passive exponents ($-1/2$, $-1/3$, and $2/3$) once shear dominates over activity. The paper also finds that the zero-shear viscosity of self-avoiding active polymers is reduced by roughly a factor of two relative to passive chains, that this reduction requires excluded-volume interactions, and that zero-shear data collapse onto a universal curve as a function of $N_m Pe$.

Load-bearing premise

The model assumes a dry, overdamped polymer with no hydrodynamic interactions and with driving tangential to the bonds; if solvent-mediated flow around the chain materially changes how it aligns, shrinks, or dissipates stress, the 4/3 and 1/3 exponents may not survive in real fluids.

Editorial extensions

If this is right

  • Dilute solutions of tangentially propelled polar polymers should exhibit a viscosity drop with shear much steeper than passive solutions, with the 4/3 exponent visible in the window $10 < Wi_{Pe} < 10^3$.
  • In the same window, tumbling slows relative to passive chains, making the stretch–recoil period a mechanical signature of polar activity.
  • For large activity, the intermediate-regime response becomes independent of $Pe$, so different activities collapse onto a single master curve and activity only sets the onset shear rate.
  • The crossover to passive exponents at large $Wi_{Pe}$ means the active signature can be erased by strong flow, which matters for processing flows.

Reading between the lines

Editorial extensions of the paper, not claims the author makes directly.

  • A direct test would be to measure single-chain viscosity or tumbling of motor-driven cytoskeletal filaments in a microfluidic shear cell; slopes other than $-4/3$ and $1/3$ would signal that hydrodynamics or boundary conditions change the mechanism.
  • If full hydrodynamic simulations shift one exponent but not the other, the tight link the paper finds between $\langle G_{yy}\rangle$ and $\eta_p$ would break, suggesting the stress and conformation are less directly coupled in real solvents.
  • The same tangential-propulsion coupling might produce steepened response in extensional or oscillatory flows, but the geometric tumbling argument would need re-derivation for those kinematics.
Share X Bluesky LinkedIn Reddit HN

Editorial analysis

A structured set of objections, weighed in public.

Desk editor's note, referee report, and a circularity audit.

Referee Report

3 major / 5 minor

Summary. The manuscript uses coarse-grained Brownian dynamics simulations of a dry, overdamped, self-avoiding flexible polymer whose monomers are driven by tangential active forces, and compares the results with a Gaussian-chain analytical model developed in Ref. [49] and rederived in Appendix E. For linear shear flow, it reports that in an intermediate, activity-dominated Weissenberg-number regime the polymer gradient size shrinks as ⟨G_yy⟩/⟨G_yy^0⟩ ∼ Wi_Pe^{-4/3}, the intrinsic viscosity thins as η_p/η_p^0 ∼ Wi_Pe^{-4/3}, the alignment angle behaves as tan(2χ) ∼ Wi_Pe^{-1}, and the tumbling frequency grows only as ν_t τ_r ∼ Wi_Pe^{1/3}, before a crossover to the passive exponents (1/2 for viscosity, 2/3 for tumbling) at large Wi_Pe. The zero-shear viscosity is found to decrease with activity and to collapse as a function of N_m Pe.

Significance. If the reported exponents are correct, the paper establishes a qualitatively new scaling regime for dilute polar active polymers: tangential activity changes the rheological and dynamical exponents by large factors relative to passive chains, and the onset shear rate is controlled by Pe. The claim is concrete and falsifiable, and the manuscript includes several internal consistency checks: two independent estimators of the tumbling time agree; the main power laws are shown for N_m = 200, 400, and 1000; and the near-proportionality of η_p and ⟨G_yy⟩ over four decades in Wi is used as a cross-check. The explicit statement that hydrodynamic interactions are neglected is an appropriate caveat. The main weaknesses are that the central stress calculation has a possible origin-dependence issue and that the exponent values are quoted without quantitative uncertainty; both are fixable without changing the scope of the paper.

major comments (3)
  1. [Section VI, Eq. (E15)] The shear stress is defined as σ_xy = -∑_i ⟨(F_i^x + F_{a,i}^x) r_i^y⟩/V, and the same defining relation is used in Eq. (E15). For the active forces in this model, ∑_i F_{a,i}^x = f_a (R_N - R_1)_x/l_0, which is not zero and has a nonzero mean under shear. Consequently, the active contribution to this virial expression is not invariant under a shift of the origin of the y-coordinate unless r_i is understood to be measured relative to the polymer center of mass. The manuscript explicitly uses CM-relative coordinates for the gyration tensor in Eq. (4) and Eq. (E13), but no such statement is made for the stress. Because the central η_p/η_p^0 ∼ Wi_Pe^{-4/3} result is extracted from this σ_xy, please state the coordinate convention actually used in the simulations and recompute η_p using r_i - r_cm (or otherwise prove the origin independence of the reported values). This is a load-bearing check and should be reported explicitly.
  2. [Figs. 2(a), 5(a), 6] The paper quotes the asymptotic exponents 4/3 and 1/3 without regression intervals or any uncertainty estimate. Since the central claim is specifically that the exponents differ from the passive values 1/2 and 2/3, please include quantitative power-law fits for each Pe, with confidence intervals and the Wi_Pe ranges over which the fit is performed, and state the statistical uncertainty of the extracted exponents.
  3. [Appendix E] The analytical comparison is performed at fixed r_d = 25 for Pe = 150 rather than by solving the Lagrange multiplier μ(γ̇) self-consistently through Eqs. (E4) and (E10). The theory therefore provides a consistency check at chosen parameter values, not an independent derivation of the −4/3 exponent or the 1/3 tumbling exponent. Please state this limitation wherever the analytical model is said to 'support' or 'confirm' the simulation results.
minor comments (5)
  1. [Section V] The phrase 'aside form end effects' should be corrected to 'aside from end effects'.
  2. [Section VIII] The phrase 'suggest a root to the development' should be corrected to 'suggest a route to the development'.
  3. [Appendix E] In Eq. (E15), 'viral stress' should be 'virial stress'.
  4. [Section VI] The duplicated phrase 'for for both properties' should be reduced to 'for both properties'.
  5. [Figure 5(b)] The inset uses N_m Pe as the horizontal axis; a brief sentence in the caption explaining why this is the natural scaling variable would improve readability.

Circularity Check

1 steps flagged · score 4.0 of 10

Simulation exponents are independent, but the analytical confirmation is parameterized by a hand-set rd rather than a self-consistent solution, weakening the claimed first-principles support.

  1. fitted input called prediction [Appendix E, Eqs. (E4), (E10), final paragraph; Fig. 5(a) theory curve]
    "Analytical calculations are performed mainly for the polymer length N = 200, rd = 0, and rd = 25, where rd = 0 corresponds to the Péclet number Pe = 0 and rd = 25 corresponds to Pe = 150."

    Eq. (E18) gives the analytical shear viscosity in terms of the Lagrange multiplier µ, and Eq. (E4) is the inextensibility constraint that should determine µ. The paper does not report solving Eq. (E4); instead, it fixes rd = 25 and labels this as Pe = 150. Because Eq. (E10) defines rd = Pe/(6µ), fixing rd = 25 with Pe = 150 is the same as imposing µ = 1 by hand. The theory curves that are said to confirm the Wi^{-4/3} slope are therefore generated at a chosen value of the key parameter, so this analytical support is an input choice rather than an independent prediction of the exponent.

full rationale

The central scaling exponents are produced by direct Langevin simulations (Section II) and are not derived from or fitted to the analytical model; the Weissenberg number uses the simulation's own measured τ_r(Pe), and the zero-shear normalization η_p^0 is read from a simulated plateau, so the simulation-based claims are self-contained. The only partial-circularity issue is the analytical confirmation: App. E imports the Gaussian active-polymer framework from self-cited Ref. [49] and evaluates it at fixed rd = 25 without solving the global inextensibility constraint (E4), so the 'Pe=150 theory' curve is effectively parameterized by an imposed µ. This weakens the claim that the 4/3 and 1/3 exponents are independently predicted analytically, but it does not infect the simulation measurements. The Summary's caveat about lacking hydrodynamics is an acknowledged transferability limitation, not a circular step. The virial expression in Sec. VI/Eq. (E15) also deserves a separate well-posedness check, because Σ_i F_a_i is nonzero and lab-frame positions would make η_p origin-dependent; that is a correctness risk outside the circularity definition.

Assumptions & free parameters 2 free parameters · 3 assumptions · 0 invented entities

The central claim is supported primarily by simulation, with a supporting analytical model that contains a hand-set parameter (rd) and a self-normalization via measured η_p^0. No new physical entities are introduced. The dry-polymer assumption and the specific active-force implementation are the main domain assumptions.

free parameters (2)
  • rd = Pe/(6μ) in the analytical model = rd = 25 for Pe=150
    In App. E, the analytical curves are computed for fixed rd values (0 and 25) rather than solving the inextensibility constraint for μ self-consistently at each shear rate, so the theory's 'prediction' depends on a hand-set parameter.
  • η_p^0 (activity-dependent zero-shear viscosity) = measured from the plateau region of the same simulation data (Fig. 5b)
    The normalized shear viscosity η_p/η_p^0 uses a zero-shear value extracted from the same runs; this is a self-normalization that can hide absolute errors but does not by itself determine the scaling exponent.
assumptions (3)
  • domain assumption Dry polymer: solvent-mediated hydrodynamic interactions are neglected (Eq. (3)).
    The Langevin equation contains only local friction, conservative forces, activity, and shear; the authors acknowledge in the Summary that hydrodynamic simulations are needed for quantitative comparison with experiments.
  • domain assumption Specific tangential active force implementation, including the end-monomer forces (F_a,i = f_a(t̂_i+1 + t̂_i)/2).
    The active force at the first and last monomers is defined as f_a t̂_2/2 and f_a t̂_Nm/2, a modeling choice that affects fore-aft symmetry; Section II.
  • ad hoc to paper The analytical model in App. E uses a Gaussian (phantom) chain with a global inextensibility constraint and fixed Lagrange multiplier parameter rd.
    The theory builds on the eigenfunction expansion of Ref. [49] and fixes rd = 0 and 25 instead of computing μ(γ̇); excluded-volume is absent, which the authors cite as the reason for the passive-exponent discrepancy.

how reviews work

0 comments
Cite this review

Pith. "Pith review of Activity-enhanced shear thinning of flexible linear polar polymers." pith.science (2026). https://pith.science/paper/YEDJEYRW

@misc{pith2026250517539,
  author       = {Pith},
  title        = {Pith review of: Activity-enhanced shear thinning of flexible linear polar polymers},
  year         = {2026},
  howpublished = {\url{https://pith.science/paper/YEDJEYRW}},
  note         = {Machine review of arXiv:2505.17539}
}
read the original abstract

The rheological properties of tangentially propelled flexible polymers under linear shear flow are studied by computer simulations and are compared with analytical calculations. We find a significant impact of the coupled nonequilibrium active and shear forces on the polymer characteristics. The polar activity enhances shear-induced stretching along the flow direction, shrinkage in the transverse direction, and implies a strongly amplified shear-thinning behavior. The characteristic shear rate for the onset of these effects is determined by the activity. In the asymptotic limit of large activities, the shear-induced features become independent of activity, and for asymptotically large shear rates, shear dominates over activity with passive polymer behavior.

Figures

Figures reproduced from arXiv: 2505.17539 by the authors.

Figure 1
Figure 1. FIG. 1. Schematic of a tangentially driven active polar linear [PITH_FULL_IMAGE:figures/full_fig_p002_1.png] view at source ↗
Figure 2
Figure 2. FIG. 2. (a) Average radius-of-gyration tensor component [PITH_FULL_IMAGE:figures/full_fig_p003_2.png] view at source ↗
Figure 4
Figure 4. FIG. 4. Square root of the mean-square local distance [PITH_FULL_IMAGE:figures/full_fig_p004_4.png] view at source ↗
Figures from the paper (5 more)
Figure 6
Figure 6. Figure 6: FIG. 6. Normalized tumbling frequency [PITH_FULL_IMAGE:figures/full_fig_p005_6.png]
Figure 5
Figure 5. Figure 5: FIG. 5. (a) Normalized shear viscosity [PITH_FULL_IMAGE:figures/full_fig_p005_5.png]
Figure 8
Figure 8. Figure 8: FIG. 8. Root-mean-square local distance (R-MSLD) as a [PITH_FULL_IMAGE:figures/full_fig_p007_8.png]
Figure 9
Figure 9. Figure 9: FIG. 9. Shear viscosity, [PITH_FULL_IMAGE:figures/full_fig_p008_9.png]
Figure 10
Figure 10. Figure 10: FIG. 10. (a) Time dependence of the projection of the poly [PITH_FULL_IMAGE:figures/full_fig_p008_10.png]

Discussion (0). Continue with ORCID to comment.

Reference graph

Works this paper leans on

83 extracted references · 77 canonical work pages

  1. [49]

    Locatelli, V

    E. Locatelli, V. Bianco, and P. Malgaretti, Activity- induced collapse and arrest of active polymer rings, Phys- ical Review Letters 126, 097801 (2021)

    2021

  2. [1]

    Lauga and T

    E. Lauga and T. R. Powers, The hydrodynamics of swim- ming microorganisms, Reports on progress in physics 72, 096601 (2009)

    2009

  3. [2]

    Elgeti, R

    J. Elgeti, R. G. Winkler, and G. Gompper, Physics of mi- croswimmers—single particle motion and collective be- havior: a review, Reports on progress in physics 78, 056601 (2015)

    2015

  4. [3]

    Hatwalne, S

    Y. Hatwalne, S. Ramaswamy, M. Rao, and R. A. Simha, Rheology of active-particle suspensions, Physical review letters 92, 118101 (2004)

    work page 2004

  5. [4]

    Sokolov and I

    A. Sokolov and I. S. Aranson, Reduction of viscosity in suspension of swimming bacteria, Physical review letters 103, 148101 (2009)

    work page 2009

  6. [5]

    Gachelin, G

    J. Gachelin, G. Mino, H. Berthet, A. Lindner, A. Rous- selet, and ´E. Cl´ ement, Non-newtonian viscosity of es- cherichia coli suspensions, Physical review letters 110, 268103 (2013)

    work page 2013

  7. [6]

    H. M. L´ opez, J. Gachelin, C. Douarche, H. Auradou, and 10 E. Cl´ ement, Turning bacteria suspensions into superflu- ids, Physical review letters 115, 028301 (2015)

    work page 2015

  8. [7]

    Saintillan, Rheology of active fluids, Ann

    D. Saintillan, Rheology of active fluids, Ann. Rev. Fluid Mech. 50, 563 (2018)

    work page 2018

Show all 83 references
  1. [8]

    Kaya and H

    T. Kaya and H. Koser, Direct upstream motility in es- cherichia coli, Biophysical journal 102, 1514 (2012)

    2012

  2. [9]

    A. J. T. M. Mathijssen, N. Figueroa-Morales, G. Junot, E. Cl´ ement, A. Lindner, and A. Z¨ ottl, Oscillatory surface rheotaxis of swimming E. coli bacteria, Nat. Commun. 20, 3434 (2019)

    2019

  3. [10]

    Belitsky and G

    V. Belitsky and G. M. Sch¨ utz, Stationary RNA poly- merase fluctuations during transcription elongation, Phys. Rev. E 99, 012405 (2019)

    2019

  4. [11]

    S. C. Weber, A. J. Spakowitz, and J. A. Theriot, Non- thermal ATP-dependent fluctuations contribute to the in vivo motion of chromosomal loci, Proc. Natl. Acad. Sci. USA 109, 7338 (2012)

    2012

  5. [12]

    Zidovska, D

    A. Zidovska, D. A. Weitz, and T. J. Mitchison, Micron- scale coherence in interphase chromatin dynamics, Proc. Natl. Acad. Sci. USA 110, 15555 (2013)

    2013

  6. [13]

    Lieberman-Aiden, N

    E. Lieberman-Aiden, N. L. van Berkum, L. Williams, M. Imakaev, T. Ragoczy, A. Telling, I. Amit, B. R. La- joie, P. J. Sabo, M. O. Dorschner, R. Sandstrom, B. Bern- stein, M. A. Bender, M. Groudine, A. Gnirke, J. Stama- toyannopoulos, L. A. Mirny, E. S. Lander, and J. Dekker, C...

    2009

  7. [14]

    Smrek and K

    J. Smrek and K. Kremer, Small activity differences drive phase separation in active-passive polymer mixtures, Phys. Rev. Lett. 118, 098002 (2017)

    2017

  8. [15]

    Saintillan, M

    D. Saintillan, M. J. Shelley, and A. Zidovska, Extensile motor activity drives coherent motions in a model of in- terphase chromatin, Proc. Natl. Acad. Sci. USA 115, 11442 (2018)

    2018

  9. [16]

    Liverpool, A

    T. Liverpool, A. Maggs, and A. Ajdari, Viscoelasticity of solutions of motile polymers, Physical review letters 86, 4171 (2001)

    2001

  10. [17]

    A. F. Pegoraro, P. Janmey, and D. A. Weitz, Mechani- cal properties of the cytoskeleton and cells, Cold Spring Harbor perspectives in biology 9, a022038 (2017)

    2017

  11. [18]

    Chaudhuri, J

    O. Chaudhuri, J. Cooper-White, P. A. Janmey, D. J. Mooney, and V. B. Shenoy, Effects of extracellular ma- trix viscoelasticity on cellular behaviour, Nature584, 535 (2020)

    2020

  12. [19]

    Corominas-Murtra and N

    B. Corominas-Murtra and N. I. Petridou, Viscoelastic networks: Forming cells and tissues, Frontiers in Physics 9, 666916 (2021)

    2021

  13. [20]

    Deblais, A

    A. Deblais, A. Maggs, D. Bonn, and S. Woutersen, Phase separation by entanglement of active polymerlike worms, Physical Review Letters 124, 208006 (2020)

    2020

  14. [21]

    Here, entanglements may play a sig- nificant role, whereas our study has addressed the poly- mer properties in a dilute solution

    with increasing activity, namely a reduced shear- thinning behavior. Here, entanglements may play a sig- nificant role, whereas our study has addressed the poly- mer properties in a dilute solution. Further theoreti- cal endeavors are necessary to resolve the influence of coll...

    2020

  15. [22]

    Deblais, S

    A. Deblais, S. Woutersen, and D. Bonn, Rheology of en- tangled active polymer-like t. tubifex worms, Physical Review Letters 124, 188002 (2020)

    2020

  16. [23]

    Backholm, W

    M. Backholm, W. S. Ryu, and K. Dalnoki-Veress, Vis- coelastic properties of the nematode caenorhabditis ele- gans, a self-similar, shear-thinning worm, Proceedings of the National Academy of Sciences 110, 4528 (2013)

    2013

  17. [24]

    Malvar, B

    S. Malvar, B. S. Carmo, and F. R. Cunha, Rheology of a nematic active suspension undergoing oscillatory shear and step strain flows, Rheologica Acta 58, 771 (2019)

    2019

  18. [25]

    Z. Xu, R. Sun, W. Lu, S. Patil, J. Mays, K. S. Schweizer, and S. Cheng, Nature of steady-state fast flow in entan- gled polymer melts: chain stretching, shear thinning, and viscosity scaling, Macromolecules 55, 10737 (2022)

    2022

  19. [26]

    C. Aust, M. Kr¨ oger, and S. Hess, Structure and dynamics of dilute polymer solutions under shear flow via nonequi- librium molecular dynamics, Macromolecules 32, 5660 (1999)

    1999

  20. [27]

    P. S. Doyle, E. S. Shaqfeh, and A. P. Gast, Dynamic simulation of freely draining flexible polymers in steady linear flows, Journal of Fluid Mechanics 334, 251 (1997)

    1997

  21. [28]

    D. E. Smith, H. P. Babcock, and S. Chu, Single-polymer dynamics in steady shear flow, Science 283, 1724 (1999)

    1999

  22. [29]

    R. G. Larson, The rheology of dilute solutions of flexible polymers: Progress and problems, Journal of Rheology 49, 1 (2005)

    2005

  23. [30]

    C. M. Schroeder, R. E. Teixeira, E. S. Shaqfeh, and S. Chu, Dynamics of dna in the flow-gradient plane of steady shear flow: Observations and simulations, Macro- molecules 38, 1967 (2005)

    2005

  24. [31]

    R. G. Winkler, Semiflexible polymers in shear flow, Phys- ical review letters 97, 128301 (2006)

    2006

  25. [32]

    H.-P. Hsu, W. Paul, and K. Binder, Polymer chain stiff- ness vs. excluded volume: A monte carlo study of the crossover towards the worm-like chain model, EPL (Eu- rophysics Letters) 92, 28003 (2010)

    2010

  26. [33]

    Huang, R

    C.-C. Huang, R. G. Winkler, G. Sutmann, and G. Gomp- per, Semidilute polymer solutions at equilibrium and un- der shear flow, Macromolecules 43, 10107 (2010)

    2010

  27. [34]

    S. P. Singh, A. Chatterji, G. Gompper, and R. G. Winkler, Dynamical and rheological properties of ultra- soft colloids under shear flow, Macromolecules 46, 8026 (2013)

    2013

  28. [35]

    J. S. Hur, E. S. Shaqfeh, H. P. Babcock, D. E. Smith, and S. Chu, Dynamics of dilute and semidilute dna solutions in the start-up of shear flow, Journal of Rheology 45, 421 (2001)

    2001

  29. [36]

    R. E. Teixeira, H. P. Babcock, E. S. Shaqfeh, and S. Chu, Shear thinning and tumbling dynamics of single poly- mers in the flow-gradient plane, Macromolecules 38, 581 (2005)

    2005

  30. [37]

    Saha Dalal, A

    I. Saha Dalal, A. Albaugh, N. Hoda, and R. G. Larson, Tumbling and deformation of isolated polymer chains in shearing flow, Macromolecules 45, 9493 (2012)

    2012

  31. [38]

    R. G. Winkler, Conformational and rheological proper- ties of semiflexible polymers in shear flow, The Journal of chemical physics 133, 164905 (2010)

    2010

  32. [39]

    R. B. Bird, C. F. Curtiss, R. C. Armstrong, and O. Has- sager, Dynamics of polymeric liquids, volume 2: Kinetic theory (Wiley, 1987)

    1987

  33. [40]

    Huang, G

    C.-C. Huang, G. Gompper, and R. G. Winkler, Non- equilibrium relaxation and tumbling times of polymers in semidilute solution, Journal of Physics: Condensed Matter 24, 284131 (2012)

    2012

  34. [41]

    S. P. Singh and R. G. Winkler, Flow driven transitions of polyelectrolytes, Journal of Rheology 64, 1121 (2020)

    2020

  35. [42]

    C. M. Schroeder, R. E. Teixeira, E. S. G. Shaqfeh, and S. Chu, Dynamics of DNA in the flow-gradient plane of steady shear flow: Observations and simulations, Macro- molecules 38, 1967 (2005)

    2005

  36. [43]

    J. D. Wheeler, E. Secchi, R. Rusconi, and R. Stocker, Not just going with the flow: The effects of fluid flow on bacteria and plankton, Annual Review of Cell and Developmental Biology 35, 213 (2019)

    2019

  37. [44]

    R. E. Isele-Holder, J. Elgeti, and G. Gompper, Self- propelled worm-like filaments: spontaneous spiral for- mation, structure, and dynamics, Soft matter 11, 7181 11 (2015)

    2015

  38. [45]

    Patra, K

    P. Patra, K. Beyer, A. Jaiswal, A. Battista, K. Rohr, F. Frischknecht, and U. S. Schwarz, Collective migration reveals mechanical flexibility of malaria parasites, Nature Physics 18, 586 (2022)

    2022

  39. [46]

    Bianco, E

    V. Bianco, E. Locatelli, and P. Malgaretti, Globulelike conformation and enhanced diffusion of active polymers, Physical review letters 121, 217802 (2018)

    2018

  40. [47]

    S. K. Anand and S. P. Singh, Structure and dynamics of a self-propelled semiflexible filament, Physical Review E 98, 042501 (2018)

    2018

  41. [48]

    R. G. Winkler and G. Gompper, The physics of active polymers and filaments, The Journal of Chemical Physics 153, 040901 (2020)

    2020

  42. [50]

    C. A. Philipps, G. Gompper, and R. G. Winkler, Tangen- tially driven active polar linear polymers—an analytical study, The Journal of Chemical Physics 157 (2022)

    2022

  43. [51]

    Pincus, A

    I. Pincus, A. Rodger, and J. Ravi Prakash, Dilute poly- mer solutions under shear flow: Comprehensive quali- tative analysis using a bead-spring chain model with a fene-fraenkel spring, Journal of Rheology 67, 373 (2023)

    2023

  44. [52]

    Kaiser and H

    A. Kaiser and H. L¨ owen, Unusual swelling of a polymer in a bacterial bath, The Journal of chemical physics 141, 044903 (2014)

    2014

  45. [53]

    Eisenstecken, G

    T. Eisenstecken, G. Gompper, and R. G. Winkler, Inter- nal dynamics of semiflexible polymers with active noise, The Journal of chemical physics 146, 154903 (2017)

    2017

  46. [54]

    S. K. Anand and S. P. Singh, Conformation and dynamics of a self-avoiding active flexible polymer, Physical Review E 101, 030501 (2020)

    2020

  47. [55]

    Mart ´ ın-G´ omez, G

    A. Mart ´ ın-G´ omez, G. Gompper, and R. Winkler, Active brownian filamentous polymers under shear flow, Poly- mers 10, 837 (2018)

    2018

  48. [56]

    Panda, R

    A. Panda, R. G. Winkler, and S. P. Singh, Characteristic features of self-avoiding active brownian polymers under linear shear flow, Soft Matter 19, 8577 (2023)

    2023

  49. [57]

    R. G. Winkler and S. P. Singh, Active polar ring poly- mer in shear flow—an analytical study, The Journal of Chemical Physics 161 (2024)

    2024

  50. [58]

    Kumar, R

    S. Kumar, R. Padinhateeri, and S. Thakur, Shear flow as a tool to distinguish microscopic activities of molecu- lar machines in a chromatin loop, Soft Matter 20, 6500 (2024)

    2024

  51. [59]

    Breoni, C

    D. Breoni, C. Kurzthaler, B. Liebchen, H. L¨ owen, and S. Mandal, Giant activity-induced stress plateau in entangled polymer solutions (2023), arXiv:2310.02929 [cond-mat.soft]

    2023 arXiv

  52. [60]

    S. Das, N. Kennedy, and A. Cacciuto, The coil–globule transition in self-avoiding active polymers, Soft Matter 17, 160 (2021)

    2021

  53. [61]

    M. Das, C. F. Schmidt, and M. Murrell, Introduction to active matter, Soft Matter 16, 7185 (2020)

    2020

  54. [62]

    Fazelzadeh, E

    M. Fazelzadeh, E. Irani, Z. Mokhtari, and S. Jabbari- Farouji, Effects of inertia on conformation and dynamics of tangentially driven active filaments, Physical Review E 108, 024606 (2023)

    2023

  55. [63]

    A. R. Tejedor, J. Ram ´ ırez, and M. Ripoll, Progressive polymer deformation induced by polar activity and the influence of inertia, Phys. Rev. Res. 6, L032002 (2024)

    2024

  56. [64]

    C. M. Schroeder, R. E. Teixeira, E. S. Shaqfeh, and S. Chu, Characteristic periodic motion of polymers in shear flow, Physical review letters 95, 018301 (2005)

    2005

  57. [65]

    C. M. Schroeder, Single polymer dynamics for molecular rheology, Journal of Rheology 62, 371 (2018)

    2018

  58. [66]

    E. S. Shaqfeh, The dynamics of single-molecule dna in flow, Journal of Non-Newtonian Fluid Mechanics 130, 1 (2005)

    2005

  59. [67]

    R. M. Jendrejack, J. J. de Pablo, and M. D. Graham, Stochastic simulations of dna in flow: Dynamics and the effects of hydrodynamic interactions, The Journal of chemical physics 116, 7752 (2002)

    2002

  60. [68]

    Hsieh and R

    C.-C. Hsieh and R. G. Larson, Modeling hydrodynamic interaction in brownian dynamics: Simulations of exten- sional and shear flows of dilute solutions of high molec- ular weight polystyrene, Journal of Rheology 48, 995 (2004)

    2004

  61. [69]

    A. V. Lyulin, D. B. Adolf, and G. R. Davies, Brownian dynamics simulations of linear polymers under shear flow, The Journal of chemical physics 111, 758 (1999)

    1999

  62. [70]

    Colby, D

    R. Colby, D. Boris, W. Krause, and S. Dou, Shear thin- ning of unentangled flexible polymer liquids, Rheologica acta 46, 569 (2007)

    2007

  63. [71]

    M. Q. Tu, M. Lee, R. M. Robertson-Anderson, and C. M. Schroeder, Direct observation of ring polymer dynamics in the flow-gradient plane of shear flow, Macromolecules 53, 9406 (2020)

    2020

  64. [72]

    de Gennes, Scaling Concepts in Polymer Physics (Cornell University, Ithaca, 1979)

    P.-G. de Gennes, Scaling Concepts in Polymer Physics (Cornell University, Ithaca, 1979)

    1979

  65. [73]

    Doi and S

    M. Doi and S. F. Edwards, The theory of polymer dy- namics, Vol. 73 (oxford university press, 1988)

    1988

  66. [74]

    Panda, R

    A. Panda, R. G. Winkler, and S. P. Singh, Supplemen- tal material: Activity enhanced shear-thinning of flex- ible linear polar polymers, URL_will_be_inserted_by_ publisher (2024)

    2024

  67. [75]

    Singh, R

    S. Singh, R. K. Singh, D. Das, and S. Kumar, Kinet- ics of polymer tumbling in shear flow: A coarse-grained description, Phys. Rev. E 99, 030501 (2019)

    2019

  68. [76]

    R. Li, G. Gompper, and M. Ripoll, Tumbling and vor- ticity drift of flexible helicoidal polymers in shear flow, Macromolecules 54, 812 (2021)

    2021

  69. [77]

    P. S. Lang, B. Obermayer, and E. Frey, Dynamics of a semiflexible polymer or polymer ring in shear flow, Phys- ical Review E 89, 022606 (2014)

    2014

  70. [78]

    Deblais, K

    A. Deblais, K. Prathyusha, R. Sinaasappel, H. Tuazon, I. Tiwari, V. P. Patil, and M. S. Bhamla, Worm blobs as entangled living polymers: from topological active mat- ter to flexible soft robot collectives, Soft Matter 19, 7057 (2023)

    2023

  71. [79]

    Nguyen, Y

    C. Nguyen, Y. Ozkan-Aydin, H. Tuazon, D. I. Goldman, M. S. Bhamla, and O. Peleg, Emergent collective loco- motion in an active polymer model of entangled worm blobs, Frontiers in Physics 9, 734499 (2021)

    2021

  72. [80]

    Ozkan-Aydin, D

    Y. Ozkan-Aydin, D. I. Goldman, and M. S. Bhamla, Col- lective dynamics in entangled worm and robot blobs, Proceedings of the National Academy of Sciences 118, e2010542118 (2021)

    2021

  73. [81]

    S. S. Ding, L. J. Schumacher, A. E. Javer, R. G. Endres, and A. E. Brown, Shared behavioral mechanisms under- lie c. elegans aggregation and swarming, Elife 8, e43318 (2019)

    2019

  74. [82]

    N. Ali, S. Nand, A. Kiran, M. Mishra, and V. Mehandia, Oscillating rheological behavior of turbatrix aceti nema- todes, Physics of Fluids 35 (2023)

    2023

  75. [83]

    S. S. Saju, R. K. Manna, and P. B. S. Kumar, Dry poly- mer model for a string of pushers: Nontrivial scaling of 12 tumbling time with shear rate, Physical Review Research 6, 043126 (2024)

    2024

Pith tools

Reviewed August 7, 2026 · model on record in the stance chip above.