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arxiv: 2604.06428 · v1 · submitted 2026-04-07 · ❄️ cond-mat.soft · physics.comp-ph

Two-dimensional active polar semiflexible polymer under shear flow

Pith reviewed 2026-05-10 17:58 UTC · model grok-4.3

classification ❄️ cond-mat.soft physics.comp-ph
keywords active polymerssemiflexible filamentsshear flownegative viscositytumbling dynamicspolymer rheologymultiparticle collision dynamics
0
0 comments X

The pith

Activity makes semiflexible polymers produce negative viscosity at weak shear rates while imposing a bending-stiffness-determined scaling on their extension.

A machine-rendered reading of the paper's core claim, the machinery that carries it, and where it could break.

Numerical simulations examine how activity combined with shear flow alters the conformations, alignment, and motion of two-dimensional semiflexible polar polymers immersed in a multiparticle collision dynamics fluid. In an intermediate shear-rate window whose location depends on activity strength, the mean-square end-to-end distance measured along the flow-gradient direction follows a scaling exponent fixed by the polymer's semiflexibility. Activity is also shown to reverse the sign of the effective viscosity at low flow rates and to change the shear dependence of the polymer's tumbling period relative to both flexible active and passive cases. At the highest shear rates the flow overwhelms activity and the polymer recovers passive semiflexible behavior.

Core claim

The simulations reveal that activity in semiflexible polymers leads to conformational changes and alignment under shear, with a characteristic scaling exponent for the mean-square end-to-end distance in the gradient direction in an intermediate activity-dependent shear-rate regime, determined by the polymer's semiflexibility. The tumbling dynamics has a time scale with stronger Weissenberg number dependence than for flexible polymers. Activity impacts rheology strongly, implying negative viscosity for weak flows, while at large shear rates shear dominates and passive behavior emerges.

What carries the argument

Brownian multiparticle collision dynamics simulation of two-dimensional semiflexible active polar polymers under linear shear, with explicit bending energy and polarity.

If this is right

  • Negative viscosity at weak flows implies that the polymer suspension can accelerate rather than resist the imposed shear.
  • The observed scaling exponent for polymer extension is controlled by semiflexibility rather than by activity strength in the intermediate regime.
  • Tumbling period grows more rapidly with Weissenberg number than it does for flexible active or passive polymers.
  • At sufficiently high shear rates the system recovers the conformational statistics and rheology of passive semiflexible polymers.

Where Pith is reading between the lines

These are editorial extensions of the paper, not claims the author makes directly.

  • The two-dimensional scaling and negative-viscosity results supply a benchmark for three-dimensional filament simulations or experiments once hydrodynamic screening is accounted for.
  • Negative viscosity suggests possible microfluidic applications in which active filaments could be used to reduce effective drag at low driving rates.
  • Varying the activity implementation or the bending-energy functional would test whether the reported scaling exponent is universal for semiflexible active polymers.

Load-bearing premise

The chosen model of activity, the specific form of semiflexible bending energy, and the multiparticle collision dynamics fluid together produce results that remain valid when the same physics is realized in three dimensions or with other activity implementations.

What would settle it

Measurement of strictly positive viscosity across all weak shear rates, or absence of the semiflexibility-set scaling exponent in the intermediate regime, in either simulation or experiment.

Figures

Figures reproduced from arXiv: 2604.06428 by A. Lamura, R. G. Winkler.

Figure 1
Figure 1. Figure 1: FIG. 1. Normalized probability distribution function of the polymer [PITH_FULL_IMAGE:figures/full_fig_p003_1.png] view at source ↗
Figure 2
Figure 2. Figure 2: FIG. 2. Root mean-square end-to-end distance as a function of the [PITH_FULL_IMAGE:figures/full_fig_p003_2.png] view at source ↗
Figure 3
Figure 3. Figure 3: and the multimedia file movie2.mp4 available online for Lp/L = 0.4, Pe = 103 , Wi = 70). At very high values of the Weissenberg number, the tumbling dynamics is faster and the polymer bends completely, enhanced by activity, while slid￾ing over itself (see the multimedia file movie3.mp4 available online for Lp/L = 0.4, Pe = 103 , Wi = 103 ). The stiffer polymer ( [PITH_FULL_IMAGE:figures/full_fig_p004_3.png] view at source ↗
Figure 6
Figure 6. Figure 6: FIG. 6. Normalized probability distribution function of the polymer [PITH_FULL_IMAGE:figures/full_fig_p005_6.png] view at source ↗
Figure 7
Figure 7. Figure 7: FIG. 7. Normalized probability distribution function of the polymer [PITH_FULL_IMAGE:figures/full_fig_p005_7.png] view at source ↗
Figure 5
Figure 5. Figure 5: FIG. 5. Mean-square end-to-end distance along the gradient di [PITH_FULL_IMAGE:figures/full_fig_p005_5.png] view at source ↗
Figure 8
Figure 8. Figure 8: (a) suggest a weak logarithmic dependence of the aver￾age bond angle on WiPe for large Wi. As expected, the mean bond angles of the stiffer polymer are smaller than those of the more flexible polymer ( [PITH_FULL_IMAGE:figures/full_fig_p006_8.png] view at source ↗
Figure 9
Figure 9. Figure 9: FIG. 9. Normalized probability distribution function of the angle [PITH_FULL_IMAGE:figures/full_fig_p006_9.png] view at source ↗
Figure 10
Figure 10. Figure 10: FIG. 10. Angle [PITH_FULL_IMAGE:figures/full_fig_p007_10.png] view at source ↗
Figure 12
Figure 12. Figure 12: displays the simulation results for Lp/L = 2 and various activities. In the passive limit, the viscosity decreases approximately with the exponent −1/2 with increasing shear, as observed and predicted previously26,33,56,58,89,90. Remark￾ably, above a certain Péclet number, the viscosity is nega￾tive over an activity-dependent range of Weissenberg num￾bers, as indicted in [PITH_FULL_IMAGE:figures/full_fig… view at source ↗
read the original abstract

The nonequilibrium structural and dynamical properties of semiflexible active polar polymers subject to linear flow are studied by numerical simulations. Filaments are confined in two dimensions and immersed in a fluid described by the Brownian Multiparticle Collision Dynamics approach. The applied shear flow causes conformational changes of a polymer, aligns it along the flow direction, and induces a tumbling motion at large flow rates. In an intermediate, activity-dependent shear-rate regime, a characteristic scaling exponent for the mean-square end-to-end distance along the gradient direction is observed. This exponent appears to be determined by the semiflexibility of the polymer. The tumbling dynamics exhibits a characteristic time, with a stronger dependence on the Weissenberg number than that of flexible active or passive polymers. Activity strongly impacts the rheological properties of the semiflexible polymers, and even implies a negative viscosity for weak flows. At very large values of the shear rate, shear dominates over activity and passive-polymer behavior is assumed.

Editorial analysis

A structured set of objections, weighed in public.

Desk editor's note, referee report, simulated authors' rebuttal, and a circularity audit. Tearing a paper down is the easy half of reading it; the pith above is the substance, this is the friction.

Referee Report

3 major / 2 minor

Summary. The manuscript numerically studies the nonequilibrium conformational, dynamical, and rheological properties of two-dimensional semiflexible active polar polymers under linear shear flow, modeled with Brownian multiparticle collision dynamics (MPCD) for the fluid. Key observations include shear-induced alignment and tumbling at high rates, an activity-dependent intermediate shear-rate regime exhibiting a characteristic scaling exponent for the mean-square end-to-end distance in the gradient direction that appears controlled by the polymer's semiflexibility, a tumbling time with stronger Weissenberg-number dependence than for flexible polymers, and activity-induced negative viscosity at weak flows, with recovery of passive-polymer behavior at very high shear rates.

Significance. If the reported scaling and negative-viscosity results prove robust, they would add to the literature on active semiflexible filaments under flow, with potential relevance to biological systems such as cytoskeletal networks. The use of MPCD to incorporate hydrodynamic interactions is a methodological strength. However, the strictly two-dimensional setting and the specific polar-activity implementation limit immediate generalization to three-dimensional experimental realizations or alternative activity models.

major comments (3)
  1. [Abstract / conformational results] Abstract and conformational-results section: the statement that the scaling exponent for mean-square end-to-end distance along the gradient direction 'appears to be determined by the semiflexibility of the polymer' is presented as a numerical observation for a single bending-rigidity value; no systematic variation of the bending energy (or direct comparison to the flexible limit) is shown to isolate its effect from the activity-dependent location of the intermediate regime.
  2. [Rheology results] Rheology-results section: the claim of negative viscosity at weak flows rests on the MPCD stress-tensor evaluation under shear; the manuscript provides no explicit formula for the stress, no convergence checks with collision-cell size or particle number, and no discussion of possible 2D artifacts or sensitivity to the chosen collision rules, all of which are load-bearing for the rheological conclusion.
  3. [Methods / all results] Methods and results sections: no error bars, statistical uncertainties, or parameter table (activity strength, bending rigidity, shear rates, Weissenberg numbers) are reported for any of the scaling exponents, tumbling times, or viscosity values, preventing assessment of numerical robustness and reproducibility.
minor comments (2)
  1. [Abstract] The abstract and main text would benefit from a brief statement on how the observed scaling exponent was extracted (fitting range, number of independent runs).
  2. [Discussion / conclusions] A short discussion of the mapping (or lack thereof) from the 2D polar-activity model to three-dimensional force-dipole or non-polar realizations would clarify the scope of the conclusions.

Simulated Author's Rebuttal

3 responses · 0 unresolved

We thank the referee for the constructive comments and positive assessment of the potential significance of our work. We address each major comment below and will revise the manuscript to incorporate the suggested improvements for clarity and robustness.

read point-by-point responses
  1. Referee: [Abstract / conformational results] Abstract and conformational-results section: the statement that the scaling exponent for mean-square end-to-end distance along the gradient direction 'appears to be determined by the semiflexibility of the polymer' is presented as a numerical observation for a single bending-rigidity value; no systematic variation of the bending energy (or direct comparison to the flexible limit) is shown to isolate its effect from the activity-dependent location of the intermediate regime.

    Authors: The reported scaling exponent was obtained for the semiflexible bending rigidity employed in all simulations. While the manuscript does not include a systematic scan of bending energies, the intermediate regime is activity-dependent and the observed exponent is consistent with semiflexible behavior under shear. To isolate the role of semiflexibility, we will add a direct comparison to the flexible (zero bending rigidity) limit in the revised conformational-results section, showing that the scaling changes in the flexible case. revision: yes

  2. Referee: [Rheology results] Rheology-results section: the claim of negative viscosity at weak flows rests on the MPCD stress-tensor evaluation under shear; the manuscript provides no explicit formula for the stress, no convergence checks with collision-cell size or particle number, and no discussion of possible 2D artifacts or sensitivity to the chosen collision rules, all of which are load-bearing for the rheological conclusion.

    Authors: The stress tensor follows the standard MPCD formulation referenced in the methods, but we agree that explicit details are needed to support the negative-viscosity claim. In the revision we will insert the explicit stress-tensor expression, add convergence tests versus collision-cell size and particle number, and include a short discussion of 2D hydrodynamic artifacts and collision-rule sensitivity. revision: yes

  3. Referee: [Methods / all results] Methods and results sections: no error bars, statistical uncertainties, or parameter table (activity strength, bending rigidity, shear rates, Weissenberg numbers) are reported for any of the scaling exponents, tumbling times, or viscosity values, preventing assessment of numerical robustness and reproducibility.

    Authors: We concur that error bars and a parameter summary are essential for reproducibility. The revised manuscript will contain a table listing all key parameters (activity strength, bending rigidity, shear rates, Weissenberg numbers) and will display error bars on all reported quantities, obtained from ensemble averages over multiple independent runs. revision: yes

Circularity Check

0 steps flagged

No circularity: results are direct outputs of numerical simulations with no derivation chain reducing to inputs or self-citations.

full rationale

The manuscript relies entirely on Brownian MPCD simulations of a 2D semiflexible active polar polymer model under shear. All reported quantities (conformational changes, scaling of mean-square end-to-end distance, tumbling times, and viscosity including negative values) are measured observables from the simulations. No analytical derivation, fitted-parameter prediction, or load-bearing self-citation is invoked to obtain the central claims; the scaling exponent is stated as an observed numerical feature whose origin is attributed to semiflexibility within the chosen model. Self-citations, if present, are not used to justify uniqueness or to close any loop. The work is therefore self-contained against external benchmarks and receives the default non-circularity score.

Axiom & Free-Parameter Ledger

3 free parameters · 1 axioms · 0 invented entities

The central claims rest on a standard numerical method (Brownian MPCD) plus model choices for activity and bending stiffness; no new entities are postulated.

free parameters (3)
  • activity strength
    Parameter controlling internal driving force, varied to define activity-dependent regimes.
  • bending rigidity
    Controls semiflexibility, directly tied to the reported scaling exponent.
  • shear rate
    External control parameter scanned across regimes.
axioms (1)
  • domain assumption Brownian Multiparticle Collision Dynamics correctly captures the hydrodynamic interactions between polymer and solvent in 2D.
    Invoked as the fluid solver without further justification in the abstract.

pith-pipeline@v0.9.0 · 5459 in / 1334 out tokens · 24414 ms · 2026-05-10T17:58:01.121992+00:00 · methodology

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Lean theorems connected to this paper

Citations machine-checked in the Pith Canon. Every link opens the source theorem in the public Lean library.

  • IndisputableMonolith/Cost/FunctionalEquation.lean washburn_uniqueness_aczel unclear
    ?
    unclear

    Relation between the paper passage and the cited Recognition theorem.

    In an intermediate, activity-dependent shear-rate regime, a characteristic scaling exponent for the mean-square end-to-end distance along the gradient direction is observed. This exponent appears to be determined by the semiflexibility of the polymer.

What do these tags mean?
matches
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supports
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extends
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uses
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contradicts
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unclear
Pith found a possible connection, but the passage is too broad, indirect, or ambiguous to say the theorem truly supports the claim.

Reference graph

Works this paper leans on

93 extracted references · 93 canonical work pages

  1. [1]

    author author R. G. \ Winkler \ and\ author G. Gompper ,\ title title The physics of active polymers and filaments , \ @noop journal journal J. Chem. Phys. \ volume 153 ,\ pages 040901 ( year 2020 ) NoStop

  2. [2]

    Alberts , author R

    author author B. Alberts , author R. Heald , author A. Johnson , author D. Morgan , author M. Raff , author K. Roberts ,\ and\ author P. Walter ,\ @noop title Molecular biology of the cell: seventh international student edition with registration card \ ( publisher WW Norton & Company ,\ year 2022 ) NoStop

  3. [3]

    Guthold , author X

    author author M. Guthold , author X. Zhu , author C. Rivetti , author G. Yang , author N. H. \ Thomson , author S. Kasas , author H. G. \ Hansma , author B. Smith , author P. K. \ Hansma ,\ and\ author C. Bustamante ,\ title title Direct observation of one-dimensional diffusion and transcription by E scherichia coli RNA polymerase , \ https://doi.org/http...

  4. [4]

    author author Y. X. \ Mejia , author E. Nudler ,\ and\ author C. Bustamante ,\ title title Trigger loop folding determines transcription rate of E scherichia coli's RNA polymerase , \ https://doi.org/10.1073/pnas.1421067112 journal journal Proc. Natl. Acad. Sci. USA \ volume 112 ,\ pages 743 ( year 2015 ) NoStop

  5. [5]

    Di Pierro , author D

    author author M. Di Pierro , author D. A. \ Potoyan , author P. G. \ Wolynes ,\ and\ author J. N. \ Onuchic ,\ title title Anomalous diffusion, spatial coherence, and viscoelasticity from the energy landscape of human chromosomes , \ https://doi.org/10.1073/pnas.1806297115 journal journal Proc. Natl. Acad. Sci. USA \ volume 115 ,\ pages 7753 ( year 2018 ) NoStop

  6. [6]

    Javer , author Z

    author author A. Javer , author Z. Long , author E. Nugent , author M. Grisi , author K. Siriwatwetchakul , author K. D. \ Dorfman , author P. Cicuta ,\ and\ author M. Cosentino Lagomarsino ,\ title title Short-time movement of E coli chromosomal loci depends on coordinate and subcellular localization , \ https://doi.org/10.1038/ncomms3003 journal journal...

  7. [7]

    Zidovska , author D

    author author A. Zidovska , author D. A. \ Weitz ,\ and\ author T. J. \ Mitchison ,\ title title Micron-scale coherence in interphase chromatin dynamics , \ http://www.pnas.org/content/110/39/15555.abstract journal journal Proc. Natl. Acad. Sci. USA \ volume 110 ,\ pages 15555 ( year 2013 ) NoStop

  8. [8]

    Lieberman-Aiden , author N

    author author E. Lieberman-Aiden , author N. L. \ van Berkum , author L. Williams , author M. Imakaev , author T. Ragoczy , author A. Telling , author I. Amit , author B. R. \ Lajoie , author P. J. \ Sabo , author M. O. \ Dorschner , author R. Sandstrom , author B. Bernstein , author M. A. \ Bender , author M. Groudine , author A. Gnirke , author J. Stama...

  9. [9]

    Ganai , author S

    author author N. Ganai , author S. Sengupta ,\ and\ author G. I. \ Menon ,\ title title Chromosome positioning from activity-based segregation , \ https://doi.org/10.1093/nar/gkt1417 journal journal Nucleic Acids Res. \ volume 42 ,\ pages 4145 ( year 2014 ) NoStop

  10. [10]

    Saintillan , author M

    author author D. Saintillan , author M. J. \ Shelley ,\ and\ author A. Zidovska ,\ title title Extensile motor activity drives coherent motions in a model of interphase chromatin , \ https://doi.org/10.1073/pnas.1807073115 journal journal Proc. Natl. Acad. Sci. USA \ volume 115 ,\ pages 11442 ( year 2018 ) NoStop

  11. [11]

    Goychuk, D

    author author A. Goychuk , author D. Kannan , author A. K. \ Chakraborty ,\ and\ author M. Kardar ,\ title title Polymer folding through active processes recreates features of genome organization , \ https://doi.org/10.1073/pnas.2221726120 journal journal Proc. Natl. Acad. Sci. USA \ volume 120 ,\ pages e2221726120 ( year 2023 ) NoStop

  12. [12]

    Smrek \ and\ author K

    author author J. Smrek \ and\ author K. Kremer ,\ title title Small activity differences drive phase separation in active-passive polymer mixtures , \ https://doi.org/10.1103/PhysRevLett.118.098002 journal journal Phys. Rev. Lett. \ volume 118 ,\ pages 098002 ( year 2017 ) NoStop

  13. [13]

    author author F. J. \ N \'e d \'e lec , author T. Surrey , author A. C. \ Maggs ,\ and\ author S. Leibler ,\ title title Self-organization of microtubules and motors , \ https://doi.org/10.1038/38532 journal journal Nature \ volume 389 ,\ pages 305 ( year 1997 ) NoStop

  14. [14]

    author author C. P. \ Brangwynne , author G. H. \ Koenderink , author F. C. \ MacKintosh ,\ and\ author D. A. \ Weitz ,\ title title Cytoplasmic diffusion: molecular motors mix it up , \ https://doi.org/10.1083/jcb.200806149 journal journal J. Cell. Biol. \ volume 183 ,\ pages 583 ( year 2008 ) NoStop

  15. [15]

    author author C. A. \ Weber , author R. Suzuki , author V. Schaller , author I. S. \ Aranson , author A. R. \ Bausch ,\ and\ author E. Frey ,\ title title Random bursts determine dynamics of active filaments , \ https://doi.org/10.1073/pnas.1421322112 journal journal Proc. Natl. Acad. Sci. USA \ volume 112 ,\ pages 10703 ( year 2015 ) NoStop

  16. [16]

    author author A. W. C. \ Lau , author B. D. \ Hoffman , author A. Davies , author J. C. \ Crocker ,\ and\ author T. C. \ Lubensky ,\ title title Microrheology, stress fluctuations, and active behavior of living cells , \ https://doi.org/10.1103/PhysRevLett.91.198101 journal journal Phys. Rev. Lett. \ volume 91 ,\ pages 198101 ( year 2003 ) NoStop

  17. [17]

    author author F. C. \ MacKintosh \ and\ author A. J. \ Levine ,\ title title Nonequilibrium mechanics and dynamics of motor-activated gels , \ https://doi.org/10.1103/PhysRevLett.100.018104 journal journal Phys. Rev. Lett. \ volume 100 ,\ pages 018104 ( year 2008 ) NoStop

  18. [18]

    Lu , author M

    author author W. Lu , author M. Winding , author M. Lakonishok , author J. Wildonger ,\ and\ author V. I. \ Gelfand ,\ title title Microtubule--microtubule sliding by kinesin-1 is essential for normal cytoplasmic streaming in D rosophila oocytes , \ https://doi.org/10.1073/pnas.1522424113 journal journal Proc. Natl. Acad. Sci. USA \ volume 113 ,\ pages E4...

  19. [19]

    Ravichandran , author G

    author author A. Ravichandran , author G. A. \ Vliegenthart , author G. Saggiorato , author T. Auth ,\ and\ author G. Gompper ,\ title title Enhanced dynamics of confined cytoskeletal filaments driven by asymmetric motors , \ https://doi.org/https://doi.org/10.1016/j.bpj.2017.07.016 journal journal Biophys. J. \ volume 113 ,\ pages 1121 ( year 2017 ) NoStop

  20. [20]

    Nguyen , author Y

    author author C. Nguyen , author Y. Ozkan-Aydin , author H. Tuazon , author D. I. \ Goldman , author M. S. \ Bhamla ,\ and\ author O. Peleg ,\ title title Emergent collective locomotion in an active polymer model of entangled worm blobs , \ https://doi.org/10.3389/fphy.2021.734499 journal journal Front. Phys. \ volume 9 ,\ pages 734499 ( year 2021 ) NoStop

  21. [21]

    author author D. A. \ Gagnon \ and\ author P. E. \ Arratia ,\ title title The cost of swimming in generalized newtonian fluids: experiments with C . elegans , \ https://doi.org/DOI: 10.1017/jfm.2016.420 journal journal J. Fluid Mech. \ volume 800 ,\ pages 753 ( year 2016 ) NoStop

  22. [22]

    author author S. S. \ Ding , author L. J. \ Schumacher , author A. E. \ Javer , author R. G. \ Endres , author A. Brown , author N. Barkai , author T. Mignot , author J. Hodgkin ,\ and\ author O. A. \ Igoshin ,\ title title Shared behavioral mechanisms underlie c. elegans aggregation and swarming , \ https://doi.org/10.7554/eLife.43318 journal journal eLi...

  23. [23]

    Deblais , author S

    author author A. Deblais , author S. Woutersen ,\ and\ author D. Bonn ,\ title title Rheology of entangled active polymer-like T. Tubifex worms , \ https://doi.org/10.1103/PhysRevLett.124.188002 journal journal Phys. Rev. Lett. \ volume 124 ,\ pages 188002 ( year 2020 ) NoStop

  24. [24]

    Deblais , author A

    author author A. Deblais , author A. C. \ Maggs , author D. Bonn ,\ and\ author S. Woutersen ,\ title title Phase separation by entanglement of active polymerlike worms , \ https://doi.org/10.1103/PhysRevLett.124.208006 journal journal Phys. Rev. Lett. \ volume 124 ,\ pages 208006 ( year 2020 ) NoStop

  25. [25]

    Sugi , author H

    author author T. Sugi , author H. Ito , author M. Nishimura ,\ and\ author K. H. \ Nagai ,\ title title C . elegans collectively forms dynamical networks , \ https://doi.org/10.1038/s41467-019-08537-y journal journal Nat. Commun. \ volume 10 ,\ pages 683 ( year 2019 ) NoStop

  26. [26]

    Panda , author R

    author author A. Panda , author R. G. \ Winkler ,\ and\ author S. P. \ Singh ,\ title title Activity-enhanced shear thinning of flexible linear polar polymers , \ @noop journal journal Phys. Rev. E \ volume 111 ,\ pages 055413 ( year 2025 ) NoStop

  27. [27]

    Panda , author S

    author author A. Panda , author S. P. \ Singh ,\ and\ author R. G. \ Winkler ,\ title title Analytical analysis of the conformational and rheological properties of flexible active polar linear polymers under shear flow , \ https://doi.org/10.1063/5.0307774 journal journal J. Chem. Phys. \ volume 163 ,\ pages 224905 ( year 2025 ) NoStop

  28. [28]

    Malvar , author B

    author author S. Malvar , author B. S. \ Carmo ,\ and\ author F. R. \ Cunha ,\ title title Rheology of a nematic active suspension undergoing oscillatory shear and step strain flows , \ https://doi.org/10.1007/s00397-019-01178-4 journal journal Rheol. Acta \ volume 58 ,\ pages 771--779 ( year 2019 ) NoStop

  29. [29]

    author author R. G. \ Larson ,\ @noop title The Structure and Rheology of Complex Fluids \ ( publisher Oxford University Press ,\ address New York ,\ year 1999 ) NoStop

  30. [30]

    Xu , author R

    author author Z. Xu , author R. Sun , author W. Lu , author S. Patil , author J. Mays , author K. S. \ Schweizer ,\ and\ author S. Cheng ,\ title title Nature of steady-state fast flow in entangled polymer melts: chain stretching, shear thinning, and viscosity scaling , \ @noop journal journal Macromolecules \ volume 55 ,\ pages 10737--10750 ( year 2022 ) NoStop

  31. [31]

    Mart \' n-G \'o mez , author G

    author author A. Mart \' n-G \'o mez , author G. Gompper ,\ and\ author R. G. \ Winkler ,\ title title Active B rownian filamentous polymers under shear flow , \ https://doi.org/10.3390/polym10080837 journal journal Polymers \ volume 10 ,\ pages 837 ( year 2018 ) NoStop

  32. [32]

    author author R. G. \ Winkler \ and\ author S. P. \ Singh ,\ title title Active polar ring polymer in shear flow - A n analytical study , \ @noop journal journal J. Chem. Phys. \ volume 161 ,\ pages 064902 ( year 2024 ) NoStop

  33. [33]

    Panda , author R

    author author A. Panda , author R. G. \ Winkler ,\ and\ author S. P. \ Singh ,\ title title Characteristic features of self-avoiding active B rownian polymers under linear shear flow , \ @noop journal journal Soft Matter \ volume 19 ,\ pages 8577 ( year 2023 ) NoStop

  34. [34]

    Kumar , author R

    author author S. Kumar , author R. Padinhateeri ,\ and\ author S. Thakur ,\ title title Shear flow as a tool to distinguish microscopic activities of molecular machines in a chromatin loop , \ https://doi.org/10.1039/D4SM00636D journal journal Soft Matter \ volume 20 ,\ pages 6500 ( year 2024 ) NoStop

  35. [35]

    Kaiser \ and\ author H

    author author A. Kaiser \ and\ author H. L \"o wen ,\ title title Unusual swelling of a polymer in a bacterial bath , \ https://doi.org/http://dx.doi.org/10.1063/1.4891095 journal journal J. Chem. Phys. \ volume 141 ,\ eid 044903 ( year 2014 ) NoStop

  36. [36]

    Eisenstecken , author G

    author author T. Eisenstecken , author G. Gompper ,\ and\ author R. G. \ Winkler ,\ title title Conformational properties of active semiflexible polymers , \ https://doi.org/10.3390/polym8080304 journal journal Polymers \ volume 8 ,\ pages 304 ( year 2016 ) NoStop

  37. [37]

    author author S. K. \ Anand \ and\ author S. P. \ Singh ,\ title title Structure and dynamics of a self-propelled semiflexible filament , \ @noop journal journal Phys. Rev. E \ volume 98 ,\ pages 042501 ( year 2018 ) NoStop

  38. [38]

    author author R. E. \ Isele-Holder , author J. Elgeti ,\ and\ author G. Gompper ,\ title title Self-propelled worm-like filaments: spontaneous spiral formation, structure, and dynamics , \ @noop journal journal Soft Matter \ volume 11 ,\ pages 7181 ( year 2015 ) NoStop

  39. [39]

    Bianco , author E

    author author V. Bianco , author E. Locatelli ,\ and\ author P. Malgaretti ,\ title title Globulelike conformation and enhanced diffusion of active polymers , \ https://doi.org/10.1103/PhysRevLett.121.217802 journal journal Phys. Rev. Lett. \ volume 121 ,\ pages 217802 ( year 2018 ) NoStop

  40. [40]

    author author M. S. E. \ Peterson , author M. F. \ Hagan ,\ and\ author A. Baskaran ,\ title title Statistical properties of a tangentially driven active filament , \ https://doi.org/10.1088/1742-5468/ab6097 journal journal J. Stat. Mech. Theor. Exp. \ volume 2020 ,\ pages 013216 ( year 2020 ) NoStop

  41. [41]

    author author C. A. \ Philipps , author G. Gompper ,\ and\ author R. G. \ Winkler ,\ title title Tangentially driven active polar linear polymers - an analytical study , \ @noop journal journal J. Chem. Phys. \ volume 157 ,\ pages 194904 ( year 2022 a ) NoStop

  42. [42]

    author author C. A. \ Philipps , author G. Gompper ,\ and\ author R. G. \ Winkler ,\ title title Dynamics of active polar ring polymers , \ @noop journal journal Phys. Rev. E \ volume 105 ,\ pages L062501 ( year 2022 b ) NoStop

  43. [43]

    Fazelzadeh , author E

    author author M. Fazelzadeh , author E. Irani , author Z. Mokhtari ,\ and\ author S. Jabbari-Farouji ,\ title title Effects of inertia on conformation and dynamics of tangentially driven active filaments , \ @noop journal journal Phys. Rev. E \ volume 108 ,\ pages 024606 ( year 2023 ) NoStop

  44. [44]

    author author A. R. \ Tejedor , author J. Ram \' rez ,\ and\ author M. Ripoll ,\ title title Progressive polymer deformation induced by polar activity and the influence of inertia , \ @noop journal journal Phys. Rev. Res. \ volume 6 ,\ pages L032002 ( year 2024 ) NoStop

  45. [45]

    author author R. G. \ Winkler ,\ title title Conformational properties of active polar semiflexible phantom polymers , \ @noop journal journal J. Chem. Phys. \ volume 162 ,\ pages 154903 ( year 2025 ) NoStop

  46. [46]

    Karan , author A

    author author C. Karan , author A. Chaudhuri ,\ and\ author D. Chaudhuri ,\ title title Inertia and activity: spiral transitions in semi-flexible, self-avoiding polymers , \ @noop journal journal Soft Matter \ volume 20 ,\ pages 6221--6230 ( year 2024 ) NoStop

  47. [47]

    Sanchez , author D

    author author T. Sanchez , author D. T. N. \ Chen , author S. J. \ DeCamp , author M. Heymann ,\ and\ author Z. Dogic ,\ title title Spontaneous motion in hierarchically assembled active matter , \ https://doi.org/10.1038/nature11591 journal journal Nature \ volume 491 ,\ pages 431 ( year 2012 ) NoStop

  48. [48]

    Doostmohammadi , author J

    author author A. Doostmohammadi , author J. Ign \'e s-Mullol , author J. M. \ Yeomans ,\ and\ author F. Sagu \'e s ,\ title title Active nematics , \ https://doi.org/10.1038/s41467-018-05666-8 journal journal Nat. Commun. \ volume 9 ,\ pages 3246 ( year 2018 ) NoStop

  49. [49]

    Schaller , author C

    author author V. Schaller , author C. Weber , author C. Semmrich , author E. Frey ,\ and\ author A. R. \ Bausch ,\ title title Polar patterns of driven filaments , \ https://doi.org/10.1038/nature09312 journal journal Nature \ volume 467 ,\ pages 73 ( year 2010 ) NoStop

  50. [50]

    Kawamura , author A

    author author R. Kawamura , author A. Kakugo , author K. Shikinaka , author Y. Osada ,\ and\ author J. P. \ Gong ,\ title title Ring-shaped assembly of microtubules shows preferential counterclockwise motion , \ @noop journal journal Biomacromolecules \ volume 9 ,\ pages 2277 ( year 2008 ) NoStop

  51. [51]

    Liu , author E

    author author L. Liu , author E. T \"u zel ,\ and\ author J. L. \ Ross ,\ title title Loop formation of microtubules during gliding at high density , \ @noop journal journal J. Phys.: Condens. Matter \ volume 23 ,\ pages 374104 ( year 2011 ) NoStop

  52. [52]

    author author J. J. \ Keya , author A. M. R. \ Kabir ,\ and\ author A. Kakugo ,\ title title Synchronous operation of biomolecular engines , \ @noop journal journal Biophys. Rev. \ volume 12 ,\ pages 401 ( year 2020 ) NoStop

  53. [53]

    Locatelli , author V

    author author E. Locatelli , author V. Bianco ,\ and\ author P. Malgaretti ,\ title title Active polymer rings: activity-induced collapse and dynamicl arrest , \ @noop journal journal Phys. Rev. Lett. \ volume 126 ,\ pages 097801 ( year 2021 ) NoStop

  54. [54]

    Lamura ,\ title title Excluded volume effects on tangentially driven active ring polymers , \ https://doi.org/10.1103/PhysRevE.109.054611 journal journal Phys

    author author A. Lamura ,\ title title Excluded volume effects on tangentially driven active ring polymers , \ https://doi.org/10.1103/PhysRevE.109.054611 journal journal Phys. Rev. E \ volume 109 ,\ pages 054611 ( year 2024 a ) NoStop

  55. [55]

    Janzen \ and\ author D

    author author G. Janzen \ and\ author D. A. \ Matoz-Fernandez ,\ title title Density and inertia effects on two-dimensional active semiflexible filament suspensions , \ https://doi.org/10.1039/D4SM00572D journal journal Soft Matter \ volume 20 ,\ pages 6618 ( year 2024 ) NoStop

  56. [56]

    \ Huang , author R

    author author C.-C. \ Huang , author R. G. \ Winkler , author G. Sutmann ,\ and\ author G. Gompper ,\ title title Semidilute polymer solutions at equilibrium and under shear flow , \ @noop journal journal Macromolecules \ volume 43 ,\ pages 10107 ( year 2010 ) NoStop

  57. [57]

    author author R. G. \ Winkler ,\ title title Conformational and rheological properties of semiflexible polymers in shear flow , \ @noop journal journal J. Chem. Phys. \ volume 133 ,\ pages 164905 ( year 2010 ) NoStop

  58. [58]

    author author C. M. \ Schroeder , author R. E. \ Teixeira , author E. S. G. \ Shaqfeh ,\ and\ author S. Chu ,\ title title Dynamics of DNA in the flow-gradient plane of steady shear flow: Observations and simulations , \ @noop journal journal Macromolecules \ volume 38 ,\ pages 1967 ( year 2005 a ) NoStop

  59. [59]

    Lamura \ and\ author R

    author author A. Lamura \ and\ author R. G. \ Winkler ,\ title title Semiflexible polymers under external fields confined to two dimensions , \ @noop journal journal J. Chem. Phys. \ volume 137 ,\ pages 244909 ( year 2012 ) NoStop

  60. [60]

    author author D. E. \ Smith , author H. P. \ Babcock ,\ and\ author S. Chu ,\ title title Single polymer dynamics in steady shear flow , \ @noop journal journal Science \ volume 283 ,\ pages 1724 ( year 1999 ) NoStop

  61. [61]

    author author R. E. \ Teixeira , author H. P. \ Babcock , author E. S. G. \ Shaqfeh ,\ and\ author S. Chu ,\ title title Shear thinning and tumbling dynamics of single polymers in the flow-gradient plane , \ @noop journal journal Macromolecules \ volume 38 ,\ pages 581 ( year 2005 ) NoStop

  62. [62]

    author author R. G. \ Winkler , author S. Keller ,\ and\ author J. O. \ R \"a dler ,\ title title Intramolecular dynamics of linear macromolecules by fluorescence correlation spectroscopy , \ @noop journal journal Phys. Rev. E \ volume 73 ,\ pages 041919 ( year 2006 ) NoStop

  63. [63]

    author author C. M. \ Schroeder , author R. E. \ Teixeira , author E. S. G. \ Shaqfeh ,\ and\ author S. Chu ,\ title title Characteristic periodic motion of polymers in shear flow , \ @noop journal journal Phys. Rev. Lett. \ volume 95 ,\ pages 018301 ( year 2005 b ) NoStop

  64. [64]

    Puliafito \ and\ author K

    author author A. Puliafito \ and\ author K. Turitsyn ,\ title title Numerical study of polymer tumbling in linear shear flow , \ @noop journal journal Physica D \ volume 211 ,\ pages 9 ( year 2005 ) NoStop

  65. [65]

    Pincus , author A

    author author I. Pincus , author A. Rodger ,\ and\ author J. Ravi Prakash ,\ title title Dilute polymer solutions under shear flow: Comprehensive qualitative analysis using a bead-spring chain model with a FENE-F raenkel spring , \ @noop journal journal Journal of Rheology \ volume 67 ,\ pages 373--402 ( year 2023 ) NoStop

  66. [66]

    Hatwalne , author S

    author author Y. Hatwalne , author S. Ramaswamy , author M. Rao ,\ and\ author R. A. \ Simha ,\ title title Rheology of active-particle suspensions , \ https://doi.org/10.1103/PhysRevLett.92.118101 journal journal Phys. Rev. Lett. \ volume 92 ,\ pages 118101 ( year 2004 ) NoStop

  67. [67]

    Saintillan ,\ title title The dilute rheology of swimming suspensions: A simple kinetic model , \ https://doi.org/10.1007/s11340-009-9267-0 journal journal Exp

    author author D. Saintillan ,\ title title The dilute rheology of swimming suspensions: A simple kinetic model , \ https://doi.org/10.1007/s11340-009-9267-0 journal journal Exp. Mech. \ volume 50 ,\ pages 1275 ( year 2010 ) NoStop

  68. [68]

    author author M. C. \ Marchetti , author J. F. \ Joanny , author S. Ramaswamy , author T. B. \ Liverpool , author J. Prost , author M. Rao ,\ and\ author R. A. \ Simha ,\ title title Hydrodynamics of soft active matter , \ @noop journal journal Rev. Mod. Phys. \ volume 85 ,\ pages 1143 ( year 2013 ) NoStop

  69. [69]

    Lamura \ and\ author R

    author author A. Lamura \ and\ author R. G. \ Winkler ,\ title title Tethered semiflexible polymer under large amplitude oscillatory shear , \ @noop journal journal Polymers \ volume 11 ,\ pages 737 ( year 2019 ) NoStop

  70. [70]

    Lamura , author R

    author author A. Lamura , author R. G. \ Winkler ,\ and\ author G. Gompper ,\ title title Wall-anchored semiflexible polymer under large amplitude oscillatory shear flow , \ @noop journal journal J. Chem. Phys. \ volume 154 ,\ pages 224901 ( year 2021 ) NoStop

  71. [71]

    author author M. P. \ Allen \ and\ author D. J. \ Tildesley ,\ @noop title Computer Simulation of Liquids \ ( publisher Clarendon Press ,\ address Oxford ,\ year 1987 ) NoStop

  72. [72]

    Jiang \ and\ author Z

    author author H. Jiang \ and\ author Z. Hou ,\ title title Motion transition of active filaments: rotation without hydrodynamic interactions , \ @noop journal journal Soft Matter \ volume 10 ,\ pages 1012 ( year 2014 ) NoStop

  73. [73]

    author author C. A. \ Philipps , author G. Gompper ,\ and\ author R. G. \ Winkler ,\ title title Tangentially driven active polar linear polymers - an analytical study , \ @noop journal journal J. Chem. Phys. \ volume 157 ,\ pages 194904 ( year 2022 c ) NoStop

  74. [74]

    Janzen , author J

    author author G. Janzen , author J. P. \ Miranda , author J. Mart \' n-Roca , author P. Malgaretti , author E. Locatelli , author C. Valeriani ,\ and\ author D. A. M. \ Fernandez ,\ title title Active polymer behavior in two dimensions: A comparative analysis of tangential and push--pull models , \ https://doi.org/10.1063/5.0243432 journal journal J. Chem...

  75. [75]

    Kikuchi , author C

    author author N. Kikuchi , author C. M. \ Pooley , author J. F. \ Ryder ,\ and\ author J. M. \ Yeomans ,\ title title Transport coefficients of a mesoscopic fluid dynamics model , \ @noop journal journal J. Chem. Phys. \ volume 119 ,\ pages 6388--6395 ( year 2003 ) NoStop

  76. [76]

    Ripoll , author R

    author author M. Ripoll , author R. G. \ Winkler ,\ and\ author G. Gompper ,\ title title Hydrodynamic screening of star polymers in shear flow , \ @noop journal journal Eur. Phys. J. E \ volume 23 ,\ pages 349 ( year 2007 ) NoStop

  77. [77]

    Gompper , author T

    author author G. Gompper , author T. Ihle , author D. M. \ Kroll ,\ and\ author R. G. \ Winkler ,\ title title Multi-particle collision dynamics: A particle-based mesoscale simulation approach to the hydrodynamics of complex fluids , \ @noop journal journal Adv. Polym. Sci. \ volume 221 ,\ pages 1 ( year 2009 ) NoStop

  78. [78]

    author author A. Lamura ,\ title title Tethered flexible polymer under oscillatory linear flow , \ https://doi.org/https://doi.org/10.1016/j.apnum.2024.07.009 journal journal Applied Numerical Mathematics \ volume 205 ,\ pages 206--214 ( year 2024 b ) NoStop

  79. [79]

    LeDuc , author C

    author author P. LeDuc , author C. Haber , author G. Boa ,\ and\ author D. Wirtz ,\ title title Dynamics of individual flexible polymers in a shear flow , \ @noop journal journal Nature \ volume 399 ,\ pages 564 ( year 1999 ) NoStop

  80. [80]

    author author J. F. \ Ryder \ and\ author J. M. \ Yeomans ,\ title title Shear thinning in dilute polymer solutions , \ @noop journal journal J. Chem. Phys. \ volume 125 ,\ pages 194906 ( year 2006 ) NoStop

Showing first 80 references.